Any use of images of this site must be accompanied with a reference to the author

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HESPERIIDAE (64 pictures) Last updated in August 2014

PAPILIONIDAE (71 pictures) Last updated in August 2014

PIERIDAE (71 pictures) Last updated in August 2014

LYCAENIDAE (+RIODINIDAE) (165 pictures) Last updated in May 2009

NYMPHALIDAE Last updated in August 2014

                  Apaturinae+Limenitinae+Nymphalinae (65 pictures)

                  Heliconiinae (91 pictures)

                  Melitaeinae (61 pictures)

                  Elymniini + Melanargiini (31 pictures)

                  Coenonymphini + Erebiini + Maniolini (Satyrinae) (109 pictures)

                  Oeneini + Satyrini (60 pictures)

                  Lybitheinae (1 picture)

MOTHS (12 pictures)

EXOTIC LEPIDOPTERA (2113 pictures)  Last updated in November 2019

          

Here is my article Advices of a Possessed One in a Novosibirsk photographic journal "Photo. Siberian Succiess" about how to photograph insects, especially butterflies, which can be found (in Russian only) in the reference given.

For particular information on the butterflies of Ural, Siberia, and the Russian Far East see my English translation of the book Y. Korshunov, P. Gorbunov. [Butterflies of the Asian Part of Russia. A Handbook]. The Ural University Press, Ekatergiburg - 1995 (in Russian)

Visit a page of my lepidopterological friend and teacher, Dr. Vladimir Dubatolov, with some of his more than 200 publications available online.

Kosterin, O. E. 1987. O bulavousykh cheshuekrylykh khrebta Katunskie 
Belki (Altai) [On club-antenned lepidopterans of the Katunskie Belki 
Range (Altai)]. - In: Bulavousye cheshuekrylye SSSR [Club-antenned 
Lepidopterans of the USSR], Abstracts of reports to the seminar 
"Systematics, faunistics, ecology and protection of butterflies", 
2-5 October 1987, Novosibirsk - P. 56-59. (in Russian)

Kosterin O.E. Dnevnye cheshuekrylye (Lepidoptera, Diurna) krebta Katunskie
Belki (Tsentral'nyi Altai). [Butterflies (Lepidoptera, Diurna) of the Katunskie 
Belki mountain range (Central Altai)]. Deposited in VINITI - 1993 - 278-B93 - 46 p.

Dubatolov, V.V., Antonova, E.M., Kosterin, O.E. Eversmannia exornata 
(Eversmann, 1837) - the only known representative of the Epiplemidae 
family in West Palearctic. Actias. Russian Journal of Scientific 
Lepidopterology - 1994 - Vol. 1 - Iss. 1-2 - P. 19-23.

Kosterin O.E. Butterflies (Lepidoptera, Diurna) of the Katunskii 
mountain ridge (Central Altai). Actias. Russian Journal of Scientific 
Lepidopterology - 1994 - Vol. 1 - Iss. 1-2 - P. 45-76.

Kosterin O.E. Butterflies (Lepidoptera, Diurna) of the Koni Peninsula 
(Magadan region). Actias. Russian Journal of Scientific Lepidopterology 
- 1994 - Vol. 1 - Iss. 1-2 - P. 77- 81. 

Dubatolov, V.V., Kosterin, O.E. A new subspecies of the Euchloe 
ausonia Hunber species group from the highlands of the Dzhungarian 
Alatau (East Kazakhstan). Atalanta - 1994 - Vol. 25 (3/4) - P. 513-520. 

Korshunov, Y.P., O.E. Kosterin. Dnevnye cheshuekrylye (Lepidoptera, Diurna),
predlagaemye dlya vkkyucheniya v Krasnuyu Knigu Respubliki Altai.
[Butterflies (Lepidoptera, Diurna) proposed for inclusion to the Red Book
of Altai Republic]. In: Materialy k Krasnoi knige Respubliki Altai (zhivotnye).
[Materials for the Red Data Book of Altai Republic (animals)], N. P. Malkov, ed. 
- Gorno-Altaisk State University - Gorno-Altaisk - 1995 - p. 96-97 (in Russian)

Korshunov, Y.P., O.E. Kosterin, G.S. Zolotarenko, V.V. Dubatolov, R.Yu. Vodyanov,
Yu. P. Malkov. Materialy po cheshuekrylym (Lepidoptera, Insecta) k
Krasnoy Knige Respubliki Altai. [Materials on Lepidoptera (Insecta) for 
the Red Data Book of Altai Republic]. In: Materialy k Krasnoi knige respubliki 
Altai (zhivotnye). [Materials for the Red Book of Altai Republic (animals)], 
N. P. Malkov, ed. - Gorno-Altaisk State University - Gorno-Altaisk - 1995 - 
p. 97-115 (in Russian)

Korshunov, Yu.P., Kosterin O.E. Items: Driopa clarius (Eversmann, 1843);
Erebia kindermanni Staudinger, 1881. In: The Red Book of Altai Republic.
Animals. Novosibirsk. 1996. pp. 44-49, 54-56. (in Russian) 

Dubatolov, V.V., Yu.P. Korshunov, P.Yu. Gorbunov, O.E. Kosterin, A.L. Lvovsky.
A review of the Erebia ligea-complex (Lepidoptera, Satyridae) from
Eastern Asia. Trans. lepid. Soc. Japan. - 1998 - Vol. 49 - P. 177-193.

Dubatolov, V.V., O.E. Kosterin. Istoriya i proiskhozhdenie nemoral'noi fauny
cheshuekrylykh v Sibiri [The history and oridin of the fauna of nemoral
Lepidoptera in Siberia]. In: Biologicheskoe raznoobrazie zhivotnykh Sibiri
[Bilogical Diversity of Animals in Siberia], materials of scientific
conference devoted to 110 anniversary of the beginning of regular zoological
studies in Siberia. Tomsk, 28-30th of October 1998. P. 50-52. (in Russian) 
Available in Russian and English

Kosterin, O.E. K faune dnevnykh babochek (Lepidoptera, Diurna) goroda Omska
i ego blizhaishikh okrestnostei. [To the butterfly (Lepidoptera, Diurna) fauna
of the Omsk city and its closest surroundings]. In: Biologicheskoe raznoobrazie
zhivotnykh Sibiri [Bilogical Diversity of Animals in Siberia], materials
of scientific conference devoted to 110 anniversary of the beginning
of regular zoological studies in Siberia. Tomsk, 28-30th of
October 1998. P. 69-70. (in Russian)

Dubatolov, V.V., O.E. Kosterin. Butterflies (Lepidoptera, Hesperioidea,
Papilionoidea) of the Dauria International Nature Reserve]. In: Nasekomye
Daurii i sopredel'nykh territorii (Sbornik nauchnykh trudov). Vypusk II.
[Insects of Dauria and Adjacent Terrtories. (Collected scientific works).
Issue II]. - 1999 - Novosibirsk - P. 138-194. (in Russian, Engl. res.)

Dubatolov, V.V., O.E. Kosterin. Butterflies (Lepidoptera, Hesperioidea,
Papilionoidea) of the Argun' basin]. In: Nasekomye Daurii i sopredel'nykh
territorii (Sbornik nauchnykh trudov). Vypusk II. [Insects of Dauria and
Adjacent Terrtories. (Collected scientific works). Issue II]. - 1999 -
Novosibirsk - P. 195-221. (in Russian)
English translation available here

Kosterin, O. E. 2000. Description of a new subspecies of the
Boloria pales (Denis et Schiffermuller, 1775) sensu Warren (1944)
(Lepidoptera, Nymphalidae) from Altai. - Far East Entomologist, No. 86 - P. 1-10.

Ivonin, V. V., Kosterin, O. E. 2000. A new subspecies of
Polyommatus icadius Groun-Grshimailo, 1890) from the Russian
Altai. - Atalanta Vol. 31 - No 1/2 - p. 171-177, colour plate XIII.

Dubatolov, V.V., Kosterin, O.E. 2000. Nemoral species of Lepidoptera
(Insecta) in Siberia: a novel view on their history and the timing of
their disjunctions. - Entomologica Fennica - Vol. 11 - p. 141-166.

Krasnaya Kniga Chitinskoi oblasti i Aginskogo Buryatskogo Avtonomnogo
okruga. Zhivotnye [Red Book of Chita Province and Aginskii Buryat Autonomous
Region. Animals.]. A. M. Vozmilov, ed. - 2000 - Poisk - Chita. Items:
by O. E. Kosterin: Strelka v-chernoe, Cercion v-nigrum (Needham, 1930) -
p. 140-141; Dedka Maaka, Anisogomphus maackii (Selys, 1872) [= Gomphus m-flavum
(Selys, 1872)] - p. 141-142; Dozorshchik iyul'skii, Anax parthenope Selys,
1839, ssp. julius Brauer, 1866 - p. 142-143; Brodyazhka ryzhaya, Pantala
flavescens (Fabricius, 1798) - p. 145-146. By O. E. Kosterin and E. A.
Maksimenko: Babka Zal'berga - Somatochlora sahlbergi Trybom, 1889 - p. 143-144.
By V. V. Dubatolov and O. E. Kosterin: Golubyanka Kiana - Plbejidea cyane
(Eversmann, 1837) - p. 186. (in Russian)

Krasnaya Kniga Novosibirskoi oblasti. Zhivotnye [Red data Book of 
Novosibirsk Province. Animals]. - 2000 - Goskomekologiya
Novosibirskoi Oblasti - Novosibirsk - 316 p. Items: by O. E. Kosterin: 
Parusnik Shtubbendorfa, Parnassius stubbendorffii (Menetries,
1849) - p. 247-248. O. E. Kosterin, V. V. Dubatolov: Perlamutrovka
neparnaya reliktovaya, Damora sagana relicta Korshunov, 1984 - p. 255-256.
O. E. Kosterin, O. G. Berezina: Chernushka eniseiskaya -Erebia jeniseiensis
(Trybom, 1877) - p. 257-258. O. E. Kosterin, V. V. Ivonin: Chernushka
Tsiklop, Erebia cyclopia (Eversmann, 1844) - p. 259-260. V. V. Dubatolov, 
O. E. Kosterin: Shashechnista Shtandelya, Melitaea plotina standeli Dubatolov, 
1997 - p. 251-252; Brazhnik Prozerpina, Proserpinus proserpina (Pallas, 1772) 
- p. 269-270; Pavlinii glaz malyi nochnoi, Eudia pavonia (Linnaeus, 1758) - 
p. 271-272; Epiplema ukrashennaya, Eversmannia exornata (Eversmann, 1837) - 
p. 273. V. V. Dubatolov, O. E. Kosterin, V. V. Ivonin: Apollon obyknovennyi, 
Parnasssius apollo (Linnaeus, 1758) - ‘. 245-246.

Kosterin, O. E. 2002 New records of butterflies
(Lepidoptera, Diurna) or the Tyva Republic [Tuva]. III. Hitherto not
reported species and some considerations about the Erebia magdalena
Strecker, 1880 and Polyomatus icarus (Rottemburg, 1775) groups.
- Atalanta -  Vol. 33 - No. 1/2 - p. 177-187, pl. 8-9.

Kosterin, O. E., Ponomarev, K. B. 2002. New data on the fauna of butterflies (Lepidoptera, Diurna)
of the city of Omsk and its closest surroundings. - Euroasian Entomological
Journal - Vol. 1 - Issue 1 - P. 111-114. (in Russian).

Krasnaya kniga Respubliki Tyva. Zhivotnye [Red Data Book of Tyva Republic.
Animals, N. I. Putuintsev, ed.]. 2002. SO RAN Press, Geo Branch, Novosibisrk,
167 p. Articles: Kosterin, O. E. Chernushka Erinnin (sayanskaya) Erebia 
erynnin Warren, 1932 (syn. E. sajanensis Korshunovm 1972) - p. 41; Kosterin, 
O. E. Priidak N. V. Barkhatnitsa Hyubnera (barkhatnitsa Narika) Hyponephele 
huebneri Kocak, 1980 (syn. H. narica Huebner, 1808-1813) - p. 40. (in Russian)

Zinchenko, V. K., Kosterin, O. E. 2002. New records of butterflies for Tyva
Republic (Tuva). III. Materials on the butterfly (Lepidoptera, Diurna)
fauna of the Todzha depression. - Zhivotnyi mir Dal'nego vostoka: Sbornik
nauchnykh trudov [Animal World of the Far East: a collected paper edition].
Blagoveshchensk State Pedagogical University - Blagoveshchensk. pp. 137-156.

Gorbunov, P., O. Kosterin. 2003. The Butterflies (Hesperioidea and 
Papilionoidea) of North Asia (Asian part of Russia) in Nature. Vol. 1. 
Rodina & Fodio and Gallery Fund, Moscow, Chelyabinsk, 392 p. 

Knyazev, S. A., O. E. Kosterin. 2003. New records of nemoral butterlies
Apatura iris (L., 1758) and Maniola jurtina (L. 1758) 
in West Siberia and their probable zoogeographical significance. 
Euroasian entomological journal 2 (3): 193-194 In Russian.

Krasnaya Kniga Omskoi oblasti. Zhivotnye, rasteniya, lishainiki [Red Data 
Book of Omsk Province. Animals, Plants, Lichens]. 2005. Omsk State Pedagogical 
University Press - Omsk, 459 p. (in Russian). Articles: O. E. Kosterin, V. V. 
Dubatolov, K. B. Ponomarev, I. I. Bogdanov. Parnassius apollo (Linnaeus, 1758) 
(42-43); O. E. Kosterin, K. B. Ponomarev. Zegris eupheme (Esper, 1782) (p. 43-44); 
O. E. Kosterin. Ahlbergia frivaldszkyi (Kindermann in Lederer, 1853) (p. 44-45); 
O. E. Kosterin, I. I. Bogdanov. Apatura metis irtyshika (Korshunov, 1982) (p. 45-47); 
O. E. Kosterin, S. A. Kyazev. Apatura iris (Linnaeus, 1758) (p. 47-48)

Kosterin, O. E. 2006. O spiske nasekomykh, rekomendovannykh Orgkomitetom dlya
vvklyucheniya vo vtoroe izdanie Krasnoy Knigi Respubliki Altai [On the list
of insects recommended by the Organising Commettee for inclusion into the second
edition of the Red Data Book of Altai Republic]. In:  Redkie zhivotnye
Respubliki Altai. Materialy po podgotovke vtorogo izdaniya Krasnoy Knigi
Respubliki Altai [Rare Animals of Latai Republic. Materials on preparation 
of the second edition of the Red Data Book of Altai Republic, N. P. Malkov, ed.,
Gorno-Altaiskiy Pedagogical University, Gorno-Altaisk, pp. 11-16.

Ivonin, V. V., O. E. Kosterin, A. Yu. Haritonov, R. V. Yakovlev. 2006. 
Predvaritel'nye ocherki po nasekomm dlya Krasnoy Knigi Respubliki Altai 
[Preliminary sketches on insects for the Red Data Book of Altai Republic]. 
In:  Redkie zhivotnye Respubliki Altai. Materialy po podgotovke vtorogo 
izdaniya Krasnoy Knigi Respubliki Altai [Rare Animals of Latai Republic. 
Materials on preparation of the second edition of the Red Data Book of 
Altai Republic, N. P. Malkov, ed., Gorno-Altaiskiy Pedagogical University, 
Gorno-Altaisk, pp. 21-58.

Gorbunov, P., Kosterin, O., 2007. The Butterflies (Hesperioidea and 
Papilionoidea) of North Asia (Asian part of Russia) in Nature. Vol. 2. 
Rodina & Fodio, Moscow. 408 pp.

Kosterin, O. E. New records of butterflies (Lepidoptera: Hesperioidea
et Papilionoidea) for Tyva Republic (Tuva). IV. The early summer 
aspect of the fauna of the protected zone of the Khan-Deer cluster of the
Ubsunurskaya Kotlovina State Biosphaeral Nature Reserve (West Sayan). 
Eversmannia. Entomological studies in Russia and adjacent regions 10: 50-58
(in Russian)

Kosterin, O. E., S. A. Knyazev, A. A. Poteiko, K. B. Ponomarev, 
T. F. Kosheleva, V. Yu. Teploukhov. 2007. New records of 
butterflies (Lepidoptera, Rhopalocera) in Omskaya and Tomskaya oblast'. 
Eurasian Entomological Journal 6 (4): 473-482. (in Russian). 

Kosterin O. E. 2007. New data on butterfly fauna 
(Lepidoptera, Diurna) of the Katunskii mountain range (Central Altai). 
Altaiskii Zoologicheskikii zhurnal [Altaian Zoological Journal]. Issue 1, 
p. 24-34.

Kosterin, O. E., M. G. Sergeev, V. V. Dubatolov. 2007. Dnevnye babochki 
(Lepidoptera, Diurna) Akademgorodka [Butterflies (Lepidoptera, Diurna) 
of Akademgorodok]. In: Priroda Akademgorodka: 50 let spustya [The Nature 
of Akademgorodok: 50 years passed], I. F. Zhimulev (ed.), SO RAN  Publishing 
House, Novosibirsk, pp. 105-133. (In Russian)

Sergeev M. G., O. E. Kosterin. 2007. Golubyanki roda Maculinea v Novosibirskom
Akademgorodke [Blues of the genus Maculinea in Novosibirsk Academy Town]. In: 
Priroda Akademgorodka: 50 let spustya [The Nature of Akademgorodok: 50 years passed], 
I. F. Zhimulev  (ed.), SO RAN  Publishing House, Novosibirsk, pp. 134-139. (In Russian)

Kosterin O. E., V. V. Dubatolov. 2007. Lokal'naya populyatsiya redkogo
okhranyaemogo vida babochek Eversmannii ukrashennoi (Eversmannia exornata
Eversmann, 1837) (Epiplemidae, Lepidoptera) v Akademgorodke  [The local population
of a rare protected moth species Eversmannia exornata (Eversmann, 1837) 
(Epiplemidae, Lepidoptera) in Akademgorodok]. In:  Priroda Akademgorodka: 50 let 
spustya [The Nature of Akademgorodok: 50 years passed], I. F. Zhimulev  (ed.), 
SO RAN  Publishing House, Novosibirsk, pp. 140-144. (In Russian)

Krasnaya Kniga Respubliki Altai. Zhivotnye [Red Data Book of Altai
Republic. Animals]. 2007. Docent N. P. Malkov, ed., Gorno-Altaisk. Articles:
V. V. Dubatolov, O. E. Kosterin: Eversmannia exornata (Eversmann, 
1837) (p. 101-102); O. E. Kosterin: Sympetrum croceolum Selys, 1883 
(p. 47-50); Anax parthenope Selys, 1839 (p. 50-52); Ischnura 
pumilio Charpentier, 1825 (p. 52); Nechalennia speciosa Charpentier, 
1840 (á. 53-56); Neolycaena davidii (Oberthur, 1881) (p. 93-96); 
O. E. Kosterin, V. V. Ivonin: Polyommatus icadius (Groum-Grshimailo, 1890) 
(p. 98-101); O, E. Kosterin, R. V. Yakovlev: Clossiana matveevi P. Gorbunov 
et Korshunov, 1995 (p. 74-76); Hyponephele cadusina (Staudinger, 1881) 
(p. 88-90); O. E. Kosterin, R. V. Yakovlev, A. V. Bondarenko: Erebia embla 
(Thunberg, 1891) (p. 77-79); O. E. Koserin, R. V. Yakovlev, M. A. Lebedeva:
Erebia kindermanni Staudinger, 1881 (p. 81-83); O. E. Kosterin, R. V.
Yakovlev, P. Y. Malkov: Neolycaena falkovitchi Zhdanko et Korshunov, 1984 
(p. 92-93); O. E. Kosterin, R. V. Yakovlev, P. Y. Malkov, Y. P. Malkov, 
A. V. Bondarenko: Parnassius clarius (Lederer, 1853). (= P. ariadne 
(Eversmann, 1843) (p. 66-69); P. Y. Malkov, R. V. Yakovlev, O. E. Kosterin, 
Y. P. Malkov, A. V. Bondarenko: Damora sagana (Doubleday, 1847) (p. 71-73); 
R. V. Yakovlev, O. E. Kosterin: Parnassius tenedius Eversmann, 1851 
(p. 69-71); Glaucopsyche argali (Elwes, 1899) (p. 96-97); 
Erebia disa (Thunberg, 1891) (p. 79-80); Erebia fletcheri 
Elwes, 1899 (p. 83-84); Oeneis elwesi Staudinger, 1901 (p. 84-86); 
Oeneis aktashi Lukhtanov, 1984 (p. 86-88); R. V. Yakovlev, 
O. E. Kosterin, P. Y. Malkov, Y. P. Malkov, A. V. Bondarenko, M. A. Lebedeva, 
S. V. Chesnokova, O. P. Vozniichuk: Parnassius apollo (Linnaeus, 1758) 
(p. 63-66); A. Y. Haritonov, O. E. Kosterin: Macromia amphigena 
fraenata Martin, 1906 (p. 56-58) (In Russian).

Kosterin O. E. 2008. Records of Erebia kefersteinii (Satyridae, Lepidoptera) 
at moderate elevations in West Altai and Kuznetskii Alatau. Altai Zoological 
Journal 2: 95-97 (in Russian)

Krasnaya Kinga Novosibirskoi oblasti [Red Data Book of 
Novosibirsk Province], 2nd edition, V. A. Yudkin, D. N. Shaulo (eds.), 
Arta, Novosibirsk, 2008, 528 p. (in Russian). Papers: 
Dubatolov, V. V., O. E. Kosterin - Mellicta plotina standeli Dubatolov, 1997 
- p. 65; Proserpinus proserpina (Pallas, 1772) - p. 80; Dubatolov, V. V., 
O. E. Kosterin, V. V. Ivonin - Parnassius apollo (Linnaeus, 1758) - p. 61-62;
Eudia pavonia (Linnaeus, 1758) - p. 81; Eversmannia exornata (Eversmann, 1873) 
- p. 82; Kosterin, O. E., O. G. Berezina, V. V. Ivonin - Erebia jenisseiensis 
(Trybom, 1877); Kosterin O. E., V. V. Dubatolov - Damora sagana relicta 
Korshunov, 1984; Kosterin, O. E. & V. V. Ivonin - Erebia cyclopia (Eversmann, 
1844), á. 72-73; Krasnoborov, I. M., O. E. Kosterin - Asplenium septentrionale 
(L.) Hoffm. (1753) - p. 428; Krasnoborov, I. M., O. E. Kosterin, O. G. Berezina -
Ephedra distachya L. (1753) - p. 412; Lomonosova. M. N., A. S. Erst, 
O. E. Kosterin - Ranunculus polyphyllus Waldst. et Kit. ex Willd. (1800) - 
p. 402.

Ivonin, V. V., O. E. Kosterin, S. L. Nikolaev.  2009. Butterflies (Lepidoptera, 
Diurna) of Novosibirsk Province. 1. Hesperiidae, Papilionidae, Pieridae. 
Euroasian Entomological Journal 8 (1): 85-104. (in Russian)

Kosterin, O. E. 2009. Data of 1990 and 2000 expeditions on butterfly 
(Lepidoptera, Diurna) fauna of the Ubsu-Nur intermontane hollow within 
Tyva Republic. In: Nature of Ubsu-Nur Hollow Reserve. Transaction of "Ubsunur
Hollow" State Biospheric Reserve. Issue 1, Darma Press, Krasnoyarsk, p. 212-225. 
(in Russian)

Tshikolovets, V., R. Yakovlev, O. Kosterin. The Butterflies of Altai, 
Sayans and Tuva // Series "The Butterflies of Palaearctic Asia", Vol. 7,
Published by V. Tshikolovets, Kyiv-Pardubice, 2009, 374 pp., 48 colour plates. 

Dubatolov, V.V., O.E.Kosterin. 2010. What is Matveev's Fritillary (Clossiana 
matveevi P. Gorbunov et Korshunov in Korshunov, Gorbunov, 1995)? Amurian 
zoological journal 2 (2): 184-188 (in Russian).

Rud'ko, G.K., O.E. Kosterin. 2010. Record of Meadow Brown Maniola jurtina 
(Linnaeus, 1758) (Satyriae, Lepidoptera) in Novosibirsk Province. Altai Zoological
Journal, issue 4, p. 31-32 (in Russian).

Ivonin, V.V., O.E. Kosterin, S.L. Nikolaev. 2011. Butterflies (Lepidoptera, Diurna) 
of Novosibirskaya Oblast. 2. Lycaenidae. Euroasian Entomological Journal 10 (2): 
217-242.

Ivonin, V.V., O.E. Kosterin, S.L. Nikolaev. 2011. Butterflies (Lepidoptera, Diurna) 
of Novosibirskaya Oblast, Russia. 3. Nymphalidae (without Satyrinae). Euroasian Entomological 
Journal 12 (2): 177-199. (In Russian)

Solovyev V.I., Ilinsky Y., Kosterin O.E. 2015. Genetic integrity of four species 
of Leptidea (Pieridae, Lepidoptera) as sampled in sympatry in West Siberia // Comparative 
Cytogenetics. Vol. 9. Iss. 3. P. 299-324. doi: 10.3897/CompCytogen.v9i3.4636.

Dubatolov V.V., Kosterin O.E. 2015. Is Triodia nubifer (Lepidoptera, Hepialidae) the only 
pre- or intergacial relic species of Lepidoptera in the Altai-Sayan Mountain System? 
Euroasian Entomological Journal 14 (2): 134-138.

Ivonin, V.V., O.E. Kosterin, S.L. Nikolaev. 2016. Butterflies (Lepidoptera, Diurna) 
of Novosibirskaya Oblast, Russia. 4. Nymphalidae, Satyrinae. Euroasian Entomological 
Journal 15 (2): 143-158. (In Russian)

Tshikolovets V., O. Kosterin, P. Gorbunov, R. Yakovlev. 2016. The Butterflies of Kazakhstan 
(Lepidoptera, Rhopalocera). The butterflies of Palaeaerctic Asia XI. Vadim Tshikolovets 
Publications, Pardubice, 379 pp., 57 pl.

Krasnaya Kniga Respubliki Altai. Zhivotnye [Red Data Book of Altai Republic. Animals]. 
3rd edition. 2017.A.V. Bondarenko ed., Gorno-Altaisk, 368 p. Articles: A.V. Bondarenko, 
R.V. Yakovlev, O.E. Kosterin: Erebia fletcheri Elwes, 1899 (p. 63-64); Oeneis aktashi 
Lukhtanov, 1984 (p. 66-67); V.V. Dubatolov, O.E. Kosterin: Eversmannia exornata  
(Eversmann, 1837) (p. 79); Neolycaena davidii (Oberthur, 1881) (p. 72-74); O.E. Kosterin, 
V.V. Ivonin: Polyommatus icadius (Groum-Grshimailo, 1890) (p. 76-78); O.E. Kosterin, 
R.V. Yakovlev: Hyponephele cadusina (Staudinger, 1881) (p. 68-69); O.E. Kosterin, 
M.A. Lebedeva, R.V. Yakovlev, P.Y. Malkov: Erebia kindermanni Staudinger, 1881 (p. 61-63); 
O.E. Kosterin, R.V. Yakovlev, P.Y. Malkov, Y.P. Malkov, A.V. Bondarenko: Parnassius 
clarius (Lederer, 1853). (= P. ariadne (Eversmann, 1843) (p. 51-53); P.Y. Malkov, 
O.E. Kosterin, R.V. Yakovlev: Neolycaena falkovitchi Zhdanko et Korshunov, 1984 
(p. 71-72); P.Y. Malkov, R.V. Yakovlev, O.E. Kosterin, Y.P. Malkov: Damora sagana 
(Doubleday, 1847) (p. 56-57); R.V. Yakovlev, O.E. Kosterin: Oeneis elwesi Staudinger, 
1901 (p. 65-66); R.V. Yakovlev, O.E. Kosterin, A.V. Bondarenko, Y.P. Malkov: 
Glaucopsyche argali (Elwes, 1899) (p. 74-75); R.V. Yakovlev, P.Y. Malkov, 
A.V. Bondarenko, O.E. Kosterin, Y.P. Malkov, M.A. Lebedeva, O. P. Vozniichuk: 
Parnassius apollo (Linnaeus, 1758) (p. 48-51). (In Russian.)

Ivonin, V.V., Kosterin, O.E., Nikolaev, S.N., Yudina, M.A. 2018. Butterflies 
(Lepidoptera, Diurna) of Novosibirsk Oblast’, Russia. 5. Updates and general 
discussion. Euroasian Entomological Journal. Vol. 17. Issue 1. P. 25-52.

Kosterin, O.E. 2019. Occasional photographic records of butterflies 
(Lepidoptera, Papilionoidea) in Cambodia. 1. The coastal Cardamom foothills 
(SW Cambodia), 2010-2018. // Acta Biologica Sibirica, Vol. 5, Issue 1, p. 84-105.

Kosterin, O.E. 2019. Occasional photographic records of butterflies (Lepidoptera, 
Papilionoidea) in Cambodia. 2. Ratanakiri and Mondulkiri Provinces of Eastern Cambodia, 
2013-2018. // Acta Biologica Sibirica, Vol. 5, Issue 4, p. 21-37.

Kosterin O.E. 2020. [Fauna of dragonflies (Odonata) and butterflies (Lepidoptera: 
Papilionoidea) of the Shlyuzovskoy forest-bog complex]. In N.P. Mironycheva-Tokareva 
(ed.) [Uique Nature Monument, “Skazochnyy” Shlyuzovskoy Forest-Bog Complex. 
Ecology and Protection. Materials of the regional scientifi-practical conference, 
19th April 2018]. Okarina, Novosibirsk, pp. 78-88 (in Russian)

288. Kosterin, O.E. 2019. Occasional photographic records of butterflies (Lepidoptera, 
Papilionoidea) in Cambodia. 3. Pursat, Siem Reap, Preah Vihear and Stung Treng Provinces 
in western, north-western and northern Cambodia. // Acta Biologica Sibirica. Vol. 6, 
p. 293-338

Kosterin O.E., Gabdullina, A.U., Zinshenko, V.K., 
Knyazev, S.A., Tshikolovets, V. 2022. Butterflies (Lepidoptera, Papilionoidea) of the Katon-Karagai 
National Nature Park, Kazakhstan // Trydy Katon-Karagayskogo gosudarstvennogo natsional’nogo 
prirodnogo parka [Proceedings of the Katon-Karagai State National Nature Park], Volume 2. 
Media-Alyans, Oskemen, p. 206-261.

Solovyev, V.I., Dubatolov, V.V., Vavilova, V.Y., Kosterin, O.E. 2022. Estimating range disjunction time of the Palaearctic Admirals 
(Limenitis L.) with COI and histone H1 genes. // Organisms Diversity & Evolution. V. 22. P. 975-1002. 

Gorbunov, P.Y., Kosterin, O.E. 2022. Butterflies (Lepidoptera: Papilionoidea) 
of Kamchatka in nature. // Amurian Zoological Journal, vol. 14, no. 3. Supplement, 
pp. 1–190.
 

Chartier, G., Kosterin, O.E. 2022. An annotated checklist of the butterflies 
(Lepidoptera: Papilionoidea) of Cambodia. // Camnodian Journal of Natural History, 
vol. 2022, no. 2, pp. 99–126.

Kosterin O.E. 2023. Early summer aspect of butterflies (Lepidoptera: Papilionoidea) 
of Republic of Khakassia as examined in 2000, with some additional data. Amurian Zoological 
Journal. Vol. 15, Iss. 2, p. 325-354.
 
https://doi.org/10.33910/2686-9519-2023-15-2-325-354

Kosterin, O.E., Knyazev, S.A. 2024. Plusiodonta casta (Butler, 1878) (Lepidoptera, 
Erebidae, Calpinae) found in West Sayan (Central Siberia, Russia). Acta Biologica Sibirica, 
vol. 10, pp. 1727-1732.
 https://doi.org/10.5281/zenodo.14470324