In 1995 Y. Korshunov with P. Gorbunov (Korshunov & Gorbunov, 1995) have published in Ekaterinburg, in Russian in 400 copies, a handbook of butterflies of the Asian part of Russia, which contained a lot of new taxa and some new names proposed to replace the homonyms. Unfortunately, information on these taxa is not so far available for a world-wide community of lepidopterologists. However, it is inevitable for their taxonomical work, even if they would not agree with some of these taxa. Later Y. Korshunov has published (in 200 copies Novosibirsk) the corrections and extensive new data in a special issue (Korshunov, 1996), which as well contained a number of new taxa and new names, and, in 1998, another analogous correction (Korshunov, 1998), as well with new descritions. English translations of all these descriptions are now available, which are suggested here for a kind attention of lepidopterologists. Several taxa were described in those books by other authors, and they are also included here. These are exact translations of the original descriptions. Small corrections or explanations are introduced into them in square brackets where necessary, in some cases relevant systematic notes are added below. Which taxon was published in which work can be inferred from the years of descriptions. An abbreviation "f.w.l." means "the fore wing length".
The type materials of the new taxa are kept in the collection of the Siberian Zoological Museum at the Institute of Systematics and Ecology of Animals, the Siberian Division of the Russian Academy of Sciences, Novosibirsk, partly - in the Institute of Ecology of Plants and Animals, the Ural Division of the Russian Academy of Sciences, Ekaterinburg.
Translator's notes: Translation into another language always helps to formulate the sense better. It is useful for a translator to make some suggestions to the athor to express some details in some other way to make them more precise, correct or concise. This is just the thing I was not able to afford myself when translating these original Russian descriptions. In many points I would advice the authors to formulate their notions in a more direct and precise way in Russian. (In particular, descriptions in (Korshunov, 1998) are extremely bad, to my opinion, and the corresponding taxa often seem to be very weak). Yet I had to preserve their style and terminology as much as both the foreign language as such and my notion of it allowed. I hope a reader would kindly take it into account.
Spialia orbifer pseudolugens P. Gorbunov, 1995
Original description:
"...For Siberia and Primorye the taxon lugens Staudinger,
1886, stated from the region of the Ferghana town, was reported
(de Jong, 1978). It seems, however, an independent species of no
relation to the fauna of North Asia. Butterflies from S Ural, S.
Siberia, and the mountains of E Kazakhstan are close to the
nominotypical subspecies (V. und A. Lukhtanov, 1994). From a
typical lugens from the Pamiro-Alai Mts. they differ by a dark
ground colour of the wing upperside, small size of white marks,
an even colouration of the hind wing underside (without pale-
brown spots), and also the male genitalia structure. Fro, the
subspecies orbifer they differ by more elongate wings and muddy-
greenish-brown ground colour of the wing underside (which is
usually brown in the subspecies orbifer). By these characters a
subspecies Spialia orbifer pseudolugens P. Gorbunov, sbsp. n. is
being stated. F.w.l. in the type series is 11.2-13.2 mm in males
and 13.1-13.8 in females. The hind wing underside ground colour
in females is brown-ochre, much lighter than in males.
MATERIALS: The holotype: a male - 10.06.1994, W Altai, the
station Turgusun environs, the Osinovka River valley (P.Yu.
Gorbunov). Paratypes: 9 males 1 female - the same locality; 2
males - 28.05.1995, the same locality (Yu.A. Shevnin); 2 males
- 16.06.1992, SE Altai, the village Aktash (V.G. Barkhatov); a
female - 29.06.1986, S. Ural, the station Poletaevo (A.
Razboinikov); 8 males 2 females - [the date was dropped, it is
12.06.1993] the Dzhungarian Alatau Mt. Chain, the environs of
the village Tekeli [in fact it is a city - Yu. K.], the Kora
River, 1500 m (V. Dubatolov, O. Kosterin)."
Pyrgus serratuilae shukshini Korshunov et Ivonin, 1996
Original description:
"For the mountains of S Siberia a new subspecies is being
descrobid - Pyrgus serratuilae shukshini Korshunov et Ivonin,
sbsp.n.
HOLOTYPE: a male. F.w.l. 14 mm. On the fore wing upperside the
white spots are distinct and on the hind wing they are diffuse,
greyish-white on the brownish-black background. The hind wing
underside is ground-green with white spots with distinct margins;
there is a large white oval spot at the wing base and the fore
margin, and a double rectangular spot at the cell is well
expressed. In the nominotypical subspecies the spot at the anal
angle of the hind wing is not roundish and the spot at the cell is
of an irregular shape. In [P.s.] uralensis these spots present in
some specimens, but the butterflies are small, with larger white
spots on the wing upperside. The hind wing underside has more white
scales on the ground colour.
ALLOTYPE: a female. F.w.l. 14 mm. The wing upperside is as in
the male, the hind wing is darker, almost without spots. In other
females greyish-white spots are more noticeable on the hind wing.
The patterin of the underside is as in males. The fringe in both
sexes is chequered.
The butterflies from Tuva are similar with shukshini by all the
characters, but they are smaller, f.w.l. being 11-13 mm.
MATERIALS: the holotype: a male - 12.06.1995, Altai, the Chibit
village surroundings, the Chuya River valley, a forb meadow (V.
Ivonin). The allotype: a female, the same locality and date.
Paratypes: 4 males 3 females - 12.06.1995, the same locality; a
male - 10.06.1902, the Katun' River, Ustyuba (G. Orlov); a male
- 14.08.1959, Altai, the village Kurai (L. Zheltikova), a male -
17.08.1960, teh same locality; a male - 9.06.1974, Altai, the
village Berkh-Kukuya, the Cherginskii Mt. Range (V. Dubatolov);
a male - 13.06.1989, the Novosibirsk Region, the upland [very
low] Sokur, the village Acha (Yu. Korshunov), a female -
17.06.1982, the same locality; a female - Tuva, Lake Azas (V.
Zinchenko); a male - 23.07.1988, Kyzyl (V. Zinchenko); a male
and female - 16.06 znd 3.07.1988, Tuva, 35 km west of the
village Samagaltai, theShivilig-Khem River left bank (M.
Sergeev); a male - 17.07.1990, the same locality (O. Kosterin);
a male - 1.07.1963.
The new subspecies is named by the surname of Shukshin,
Vasilii Makarovich (1929-1974), a well known Siberian writer
born in Altai."
Three new subspecies of Hesperia comma (Linnaeus, 1758)
have been described from Siberia (Korshunov & Gorbunov, 1995).
It was stated that "a variation exists in females within these
subspecies, and they have certain differences in configuration
of the teeth on the valva and other details of male
genitalia." In (Korshunov, 1996) iis stated that they differ from the
nominotypical subspecies by the ground colour, the spots, the
presence of their dark rims; in the male genitalia there are
differences in the shape of teeth on the valva. In the lateral
view the apical processus of the valva bears small teeth in ssp.
sushkini and is lower than the basal spine, in ssp. planula the
processus lacks teeth and is on the same level as the basal
spine, in ssp. lena the processus is lower than the spine and is
covered with uneven knobs.
Hesperia comma lena Korshunov et P.Gorbunov, 1995.
Original description:
"HOLOTYPE: a female. F.w.l.: 16 mm. The wing upperside is
relatively dark, brown with fulvous spots; the underside of the
hind wing and of the fore wing apex are of a muddy-green ground
colour, the inner margin of the hind wing underside being
yellowish; the white spots are contrasted but their dark rims are
not distinct as being hidden by muddy-green scales.
ALLOTYPE: a male. F.w.l.: 15 mm. On the wing upperside the
dark-fulvous colour is to a great extent displaced by dark-brown
scales. The androconial stroke contains a light stria inside. The
hind wing underside is muddy-green, with a fulvous suffusion at
the anal angle and with roundish and angular white spots, their
dark rims being masked by the scales of the ground colour.
MATERIALS: the holotype: a female, the Suntar-Khayata mountain
range, 356th km of road Khandyga-Magadan, a damp meadow on the
Suntar river bank, 19th July 1985 (V.V.Dubatolov leg.); the
allotype: a male, the Suntar-Khayata mountain range, 232-nd km
of road Khandyga-Magadan, a mountain Baranya, 1600 m altitude,
7th July 1985 (V.V.Dubatolov leg.); paratypes: male and female -
C. Yakutia, settlement Khaptagay, the Tamma river valley, 20th
July 1973 (Yu.N.Ammosov leg.); 10 males 8 females - Suntar-
Khayata mountain range, 232-nd km of road Khandyga-Magadan,
16th-27nd June and 1si-22th July 1985 (V.V.Dubatolov leg.); 2
females, Yakutsk, Botanical Garden, (V.V.Dubatolov leg.), a male
- NW. Chukotka, 100 km south of settlement Pevek, Ust'-Chaun,
24th July 1986 (V.K.Zinchenko leg.)."
Hesperia comma planula Korshunov, 1995.
Original description:
"HOLOTYPE: a male. F.w.l.: 15 mm. The fore wing apex is more
pointed than in the nominotypical subspecies; the fore wing
upperside is light-fulvous with less amount of dark scales; the
androconial stroke contains a conspicuous light streak; the hind
wing underside is yellowish-green (while in Swedish specimens it
is olive-green) with angular white spots.
ALLOTYPE: a female. F.w.l.: 16 mm. On the fore wing upperside
the yellow scales cover not all the white spots, this especially
concerns the spots at the outer margin. The hind wing underside
is yellowish-green with large white spots in dark rims.
MATERIALS: the holotype: a male, Novosibirsk region,
settlement Karasuk, Lake Krotovaya Lyaga, a birch grove edge,
29th July 1981 (V.V.Dubatolov leg.); the allotype: a female,
Novosibirsk region, 10 km NNW of town Zdvinsk, a steppefied
meadow at a road, 14th August 1990 (V.V.Dubatolov leg.);
paratypes: a female, environs of Barnaul,29th July 1902
(E.G.Rodd, leg.); 1 male 2 females - Novosibirsk region,
village Novyy Sharap, birch kolki 8th, 19th, and 28th August
1957; a female - the same locality, 2nd August 1959; Novosibirsk
region, railway station Izdrevaya, 1st July 1959; a male -
Tyumen' region, village Oktyabr'skoe, the Ob' bank, 9th July
1969 (Yu.P.Korshunov leg.); a male - surroundings of
Chelyabinsk, 30th July 1979 (A.E.Kulyginskiy leg.); a female -
Chelyabinsk region: town Troitsk surroundings, 14th August 1982
(M.F.Manapov leg.); 1 male 5 females - Novosibirsk region,
settlement Karasuk, Lake Krotovaya Lyaga, 23th July 1981; a male
- the same locality, 10th August 1990; a female - the same
locality, 11th August 1990 (V.V.Dubatolov leg.); a male -
Novosibirsk region, 10 km NNW of town Zdvinsk, 19th August 1990
(V.V.Dubatolov leg.)."
Hesperia comma sushkini Korshunov, 1995.
Original description:
"HOLOTYPE: a female. F.w.l.: 18 mm. All the light-yellow
spots are distinct on a dark ground colour of the wing
upperside; the hind wing upperside is grey-green with a light-
grey inner border; the scales at the anal angle are lighter than
others and so form a conspicuous spot; the white spots are
distinct, of intermediate size
ALLOTYPE: a male. F.w.l.: 16 mm. The wing upperside is
fulvous, the outer area of the fore wing and the margins of the
hind wing are dark; the androconial stroke contains a light
stria; the white spots of the hind wing underside are large,
distinct, with dark rims.
MATERIALS: the holotype: a female, Altai, the Chulyshman river
valley, 19th August 1980 (M.Ivonina leg.); the allotype: a male,
Altai, Katunskiy mountain range, the Nizhniy Kuragan river bank
terrace at the mouth, a steppefied meadow, 24th July 1986
(O.E.Kosterin leg.); paratypes: a female - Nizhnyaya Ustyuba,
25th July 1902 (G.Orlov leg.); 3 males - N. Pribaikalye, the
Malaya Kosa river, a larch wood edge (Velizhanin leg.); 1 male 1
female - Novosibirsk region, Bugotakskie Sopki [Bugotak Hills]
at settlement Semenovskiy, 5th August 1982 (V.V.Ivonin leg.); a
male - Krasnoyarskiy Kray, the Sayano-Shushenskiy reserve, the
Uzun-Suk river, 11-16th August 1988 (V.A.Vagin leg.); a male -
West Altai, Glubokaya, 2nd September 1990 (V.K.Zinchenko leg.).
The subspecies is named after the surname of Petr Petrovich
Sushkin (1868-1928), a known ornitologist and lepidopterologist,
who collected Lepidoptera in the mountains of South Siberia
togerther with S.S. Chetverikov."
Note: the spelling "shushkini" printed in the headline of
the original description is clearly an error, since it was clearly
stated that the taxon is named after the surnameSushkin.
Leptidea amurensis jacutia P. Gorbunov et Korshunov, 1995.
Original description:
"Butterflies from Central Yakutia differ by a noticeable
yellowish tint on the wing upperside and more intensive, than in
other subspecies, ochre suffusion on the underside of the hind
wing and on the most part, at the apex and the fore margin, of
the fore wing. By these characters we state a subspecies
Leptidea amurensis jacutia P. Gorbunov et Korshunov, sbsp.n.
MATERIALS: Holotype: a male - 2.06.1985, Yakutsk, the
botanical garden (V. Dubatolov). Paratypes: a male - 21.06.1962,
the same locality; 2 males - 28.05.1985, the same locality; a
male - 2.06.1985 - the same locality."
Pontia daplidice kulyginskyi Korshunov, 1998.
Original description:
"...Discovery of the taxon which we describe below belongs to
an Uralian colleague E. A. Kulyginsky. When sending us these miniature
butterflies he persuaded us much to pay a special attention to them.
All this took place still before discussion of the publications devoted
to the differences between daplidice and edusa. That time he stressed that
in the Chelyabinsk Province in 1975-1979 he mei with exclusively large
form of the . And only at station Bredy in 1978 in August he found
small butterflies which he at first visually confused with some light
blues.
Having received these small imagines in 1977 we found them similar with
those small specimens of which were regularly observed by us in spring
and autumn in the Novosibirsk Province. Check of these materials
nowadays have showed a difference: only the butterflies from the South Ural
are close to daplidice, and we describe them in the memory of EUvgenii
Aleksandrovich Kulyginskii as Pontia daplidice kulyginskyi Korshunov,
sbsp. n. F.w.l.: 17-20 mm. HOLOTYPE: a male, above and beneath its pattern,
as wellas that of the paratypes, little differ from edusa, but one can note
that the discal spot have a central lightening. ALLOTYPE: a female, which
also resembles edusa, but it, as well as the paratypes, has a black pattern on
the hind wing upperside more developed and has no a dark strope at ourer
margin at anal angle in front of the black rounded spots. Among the paratypes
a female is conspicuous with the fore wing length of 22 mm, in which a dark
suffusion is developed on the fore wing extending from the apical black
spots to the middle of the cell, while on the fore wing underside such a
suffusion is expressed only at the cell. The hind wing underside is also of
an aberrant nature as almost entire its surface is densely suffused with
yellowish-green scales, white spots being hardly noticeable in the center of the
wing and well expressed only along the margin.
MATERIALS: Holotype - a male, 15.08.1978, Bredy, steppe, E. A. Kulyginsky.
Holotype - a female, 15.08.1978, the same label. Paratypes - five
males and three females - 16.08.1978, a steppefied forest edge at station
Bredy. In spring 1997 on 19th May in the Orenburg Nature Reserve, in
the Burtinskaya Steppe a male with the fore wing length of 19 mm was
collected by our colleague I. Lyubechanskii. This was a first corroboration
of a reality of the subspecies described."
Euchloe ausonia dubatolovi Korshunov, 1995.
Original description:
"... The butterflies from Altai, described below, differ from
the western subspecies by a faded muddy-green hind wing
underside ground colour and smaller white spots. By these and
other traits they look like a peculiar transition (especially
in the populationf of the Chuya River valley) to E. naina.
HOLOTYPE: a male. F.w.l: 22 mm. The wing upperside is white;
the fore wing apex is suffused with dark scales, the suffusion
containing three white spots; the dark discoidal spot is
crescent-shaped. The fore wing underside is white, the dark
discoidal spot with a contrasted light discoidal vein; the apex
is suffused with dark and green scales and has the same white
spots as on the upperside; the costal margin with short black
striae. The hind wing underside is greyish-green with white spots
of different sizes, mainly with sharp edges.
ALLOTYPE: a female. F.w.l.: 23 mm. The wing upperside is
white; the dark discoidal spot is large, concave on its sides, and, as
different from E. n. naina [an error - must be E. ausonia ausonia -
Yu.K.], is not fused [along suffused veins] with the apical
suffusion, which contains three elongate white spots.
MATERIALS: the holotype: a male, Altai, Katunskiy mt. range, 15
km SW of village Katanda, interfluve of the Kuragan and Kucherla
rivers, alpine meadow, 2300 m, 13th July 1983 (V.V.Dubatolov
leg.); the allotype: a female, the same locality, 14th July 1983
(V.V.Dubatolov leg.); paratypes: Altai, Ust'-Kan district,
Karlinskiy Pass, 15th July 1970 (Kosinykh leg.); a male - the
same locality as for the holotype, 12th July 1983 (V.V.Dubatolov
leg.); 1 male 1 female - Katunskiy mt. range, the Katun' river
headwaters at the Kapchal river mouth, 1700 m, 9th July 1987
(O.Kosterin leg.); a male - the Kapchal valley, 2300 m, 10th
July 1987 (O.Kosterin leg.); 2 males 1 female - the Katun'
headwaters, stow Altyn-Bulak, an old felled land, 1800 m, 13th
July 1987 (O.Kosterin leg.); a male - Katunskiy mt. range, lower
reaches of the Koksu river, 1600 m, 2th July 1988 (O.Kosterin
leg.); a male - left bank of the Koksu river, long-forb meadow
on wood opening, 1900 m, 12th July 1988 (O.Kosterin leg.); 1
male 2 females - the Argem (Direntay) river (a tributary of the
Koksu) headwaters, an alpine meadow, 2200 m, 21th July 1988
(O.Kosterin leg.); a male, Kholzun mt. range, watershed of the
Tigirek and Khamir river headwaters, 2300 m, 17th July 1983
(V.A.Lukhtanov leg.).
The subspecies is named after the surname of Vladimir
Vladimirovich [must be Viktorovich instead of Vladimirovich -
Yu.K.], an entomologist of ISEA (Novosibirsk) who collected a
part of the type series of the new subspecies. "
Euchloe naina kuznezovi Korshunov, 1995.
Original description:
"...Below we describe the butterflies from the Yenisei River
low reaches, Taymyr, and the Putorana Plateau. They are closer
to the subspecies naina than to jacutia, but differ from the
former by in average more narrow wings and a darker hind wing
underside ground colour.
HOLOTYPE: a male. F.w.l: 22 mm. The wing upperside is white;
the fore wing costal margin from the base to the discoidal vein
is suffused with black scales, further to the apex there are
alternating black and white patches along the margin; the
discoidal vein is accompanied by a figured dark spot which is to
some extent more rounded than in E. naina naina. At the apex
there are a dark stripe 2-3 mm wide which is parallel to the
outer margin, and dark spots at the vein endings at the margin;
the space between the stripe and the margin being generally white
while in E. naina naina it is almost fully suffused. On the fore
wing underside the dark discoidal spot is larger than in E.n.
naina. The hind wing underside is almost exactly identical to
that of E.n. naina, but the veins are not so conspicuous.
ALLOTYPE: a female. F.w.l.: 23 mm. On the wing upperside the
white ground colour is almost displaced by a dark suffusion, the
white areas remaining only in the centre and at the edges of the
wing. The hind wing underside ground colour is greyish-green
without a dark suffusion; the veins are not contrasted.
MATERIALS: the holotype - the Yenisey lower reaches, the
Irbo-Keta river middle flow, 18th July 1982 (D.M.Pupavkin leg);
allotype - the same locality, 21st July 1982; paratypes: 5 males
3 females - the Yenisey lower reaches, the Rybnaya river middle
flow, 12th-21st July 1982 a male - the Yenisey lower reaches, the
Togbym river middle flow, 12th July 1982; 14 males 11 females -
the Yenisey lower reaches, the Irbo-Keta river middle flow, 18th
July 1982 (D.M.Pupavkin leg); a male - Taymyr, the Ary-Mas
forest, July 1983 (Polovinkina leg.)
The subspecies being describing was earlier reported
(Korshunov, 1985) as E. ausonia arctica, as it was designated in
the labels by Nikolay Yakovlevich Kusnezov (1873-1948), a well-
known Russian lepidopterologist and zoogeographist of the first
half of this century. "
Colias palaeno gomojunovae Koshunov, 1996.
Original description:
"...It was Kurenzov (1970) who first noticed that Chukotka is
inhabited by a peculiar variety of palaeno but he thought that
it is identical to the form of Alaska, designed by him as
schippewa (lapsus calami - Yu.K.) instead of chippewa Edw.
However, the males of chippewa resemble our specimens only by
the wing upperside, but the butterflies are much more yellow
[above] and the marginal band is wider and is not cut through by
[light] veins; the wing underside is more greenish, the discal
spots are absent or hardly noticeable only on the fore wings. In
the females from Alaska [I have studied 7 males and 3 females
from there] black discal spots are seen above and beneath, the
ground colour is yellowish. The females from Chukotka have a
yellowish-white ground colour and no discal spots. There is also
a difference in the genitalia, that was mentionned by Kurenzov
(1970). By these characters we describe a new subspecies Colias
palaeno gomojunovae Koshunov, sbsp.n.
HOLOTYPE: a male. F.w.l.20 mm. The wing upperside is yellowish
green, the marginal spots on the fore wing are wider than on the
hind wing, the colour of the marginal band is brownish, the band
is not cut through by light veins. There is no discal spot on
the fore wing upperside, but it is slightly seen on the
underside of both wings. The hind wing underside is yellowish-
green, with scattered grey scales. The fore wing underside is of
the same colour, but grey scales present only at the costal
margin. On the hind wing underside the silvery spot large, its
diameter approximately equals to the length of elongate (and
also silvery) spot on the transversal vein.
ALLOTYPE: a female. F.w.l. 21 mm. The wing upperside is
yellowish-white. The marginal band at the apex is relatively
wide, it is tapering to the anal angle, forming a kind of a
triangle. Light spots present only in its wide part. On the hind
wind there is only a narrow marginal streak at the fore angle.
The discal spots are missing. The hind wing underside is muddy-
green, a silvery spot is smaller than in the male, there is an
additional tiny round spot above it. On the fore wing upperside
there is only traces of the discal spot, the apex is of the same
colour as the hind wing, the rest part of the wing is light.
In the male paratypes the marginal band is distinctly
cut through by light veins only in 3 [of 7] specimens, which
are worn out, in fresh specimens it is darker and is covered
with silvery scales beckoming worn out later.
MATERIALS: The holotype: a male - 30.07.1968, the Magadan
Region, The Bilibino District (N. Gomoyunova); the allotype:
a female - 12.07.1968, the left bank of the Khoroshaya
River, a tributary of the Molondzha River, herbs/dwarf
birches and willows/cloudberry (N. Gomojunova). Paratypes: 2
males - 3 and 15.07.1967, the settlement Markovo on the
Anadyr' River (A.G. Mirzaeva); 2 males 1 female - 12 and
13.07.1968, the Khoroshaya River left bank; a male -
19.07.1969, the
Molondzha River floodland; 2 males 2 females - 10 and
14.07.1969, the Magadan Region, the Kegali River, a larch
parkland with the dwarf birches and willows; 2 males - 25.07
1nd 2.08.1986, West Chukotka, [the settelment] Ust'-Chaun,
100 km south of Pevek (V. Dubatolov).
The subspecies is dedicated to the memory of Nina
Petrovna Gomojunova (1933-1973) - an entomologist and
parasitologist of the Biological Institute of Siberian
Division of the Academy of Sciences, who collected in
Chukotka following the nomadic rain-deer breeders. She
tragically perished in an aircraft cetestrophe."
Colias chrysotheme elena P. Gorbunov, 1995.
Original description:
"...Below we describe very peculiar butterflies of the Altai
Mountains, the Sayans, and the mountains of Tuva, differing from
lowland subspecies by a more even and bright colouration of the
wing upperside, narrower light [streaks along the] veins in the
dark marginal band in males and darker and more contrasted
colouration of females, which tend to resemble C. thisoa by
these characters.
Colias chrysotheme elena P. Gorbunov, sbsp.n.
MALE: F.w.l. 24.4-27.9 mm (26.2 mm in the holotype). The wing
upperside is light-orange with a narrow yellowish stripe along
the fore wiong fore margin and an area of the same colour at the
anal margin of the hind wing. A slight suffusion with dark
scales is noticeable on the hind wing. The marginal band on the
fore wing in its middle part is 3.5-5 mm wide, usually all over
its length it is cut through with narrow light [streaks along
the] veins, which most often don't reach the fringe. The maximum
width of the marginal band on the hind wing is 2.4-4 mm; light
venal streaks on it are even narrower than on the fore wing, in
some specimens they are hardly seen at the internal margin of
the band. Underside the ground colour of the hind wing is
yellow, that in the central area of the fore wing is orange-
yellow, the pattern is as in the subspecies andre. The genitalia
are alike those in C. chrysotheme andre and C. thisoa [the statement
concerning thisoa is evidently of no relevance - Yu.K].
FEMALE: F.w.l. 25.5-28.7 mm (27.6 mm in the allotype). The
fore wing upperside is bright orange-red (while in the
subspecies andre they are yellwoish-orange or orange-yellow)
with a dark suffusion in the basal area and at the fore margin.
The marginal band on the fore wing is blackish-brown, 4.8-6.0 mm
wide in its narrowest part, with very contrasted 6-7 yellwo
spots. The black discal spot is large, usually of a triangular
shape. The hind wing upperside is muddy-orange (much darker than
in the females of andre) with an orange discal spot about 2 mm
in diameter. On the hind wing the marginal band contains two
yellow spots at the apex, four other yellow spots interrupt the
band fusing with the ground colour at its internal margin. The
discal spot is oval-shaped, 3.0-3.6 mm long. The wing upperside
ground colour is greenish-yellow, but orange in the central area
of the fore wing. There is a row of 3-5 black spots along the
outer margin of the wore wing. The discal spot on the fore wing
underside is somewhat larger than on the upperside and contains
a light dot. On the hind wing underside just above an oval
silvery discal spot there is an additional small silvery spot.
MATERIALS: The holotype: a male - 29.06.1993, SE Altai, the
village Aktash (V. Barkhatov). The allotype: a female -
26.07.1982, Tuva, Choon-Khem (O. Fileva). Paratypes: 3 males -
2-3.07.1971 , Tuva, the settlement Shuurmak,a steppe (S.
Nikolaev); a female - 25.06.1971, the same locality; a female -
16.07.1980, Altai Mts., the Seminskii Pass (Yu. Korshunov); 2
males - 25.06.1992, SE ALtai, the village Aktash (V. Barkhatov);
14 males 1 female - 27-29.06.1993, the same locality."
Colias thisoa nikolaevi Korshunov, 1998.
Original description:
" Specimens from the mountains of S Siberia have differences and we
describe them as
C. thisoa nikolaevi Korshunov, sbsp. n.
Holotype - a male. F.w.l. 25 mm. The wing uppertside intensive yellow
with a weak violet flash, while in the western butterflies they are
red-yellow with a stronger flash. As different from the nominotypic
subspecies the black border is somewhat narower. The silvery spot on
the hind wing underside larger.
In females fore wing upperside red-orange, brighter than in males. As
compared with other subspecies, their wings are less suffused with black
and red scales. Fore wing outer margin quite straight. The dark border
at anal angle above fore wing narrower than in the Tien Shan race
urumtsiensis Verity, 1909, without a violet tint, it contains relatively
large yellow spots of different size.
MATERIALS: holotype - a male, 20.06.1960. Altai, Kurai, the stow Tyurguno,
a steppen patch. Paratypes - five females - 14.07.1959, Chuiskaya Step',
the Yustyd' River valley at village Ak-Tal, two males - 11.07 and 27.08.1960,
Kurai, the stow Argatyt, a female and male - 20.07.1960, Kurai, a male and
three females - 7.07.1973, the valley to Aktru, 12 km south of Kurai, meadows
in front of a forest, a male - 2.07.1986, Altai Nature Reserve, Lake
Dzhulukul', 2200 m, N. Zolotukhin, a male - 20.07.1972, Tuva, Moren, a
river valley, J. Vijdalepp, three males, two females - 10.06.1992, village
Aktash, Kurai Mt. Range, a steppefied slope, 2100 m, V. Barkhatov.
The subspecies is named in the honour of Sergei L'vovich Nikolaev, who
collected many ALtai butterflies for the Zoological Museum collection and
was our faithful comrade in varuious Siberian expeditions."
GENUS RODDIA Korshunov, 1995.
Original description:
"Type species: Papilio l-album Esper, 1780.
A monotypic Holarctic genus. The imago fore wing length is 26-
32 mm. The wing shape and pattern resemble those of both
Polygonia and Nymphalis genera. The wing outer margin is
toothed. The wing upperside is reddish-fulvous with rather large
black spots and a white spot at the costal margin of each wing.
The wing bases are covered with dense soft hairs. The wing
underside is brown, the postdiscoidal area being lighter, with
dark strokes or reddish-brown.
The taxon is named in the honour of a forester and
entomologist Eugeniy Georgievich Rodd (1871-1933), an explorer
of Altai and the Upper Priobye."
Euphydryas iduna semenovi Korshunov et Ivonin, 1996
Original description:
"The upland Kuznetskoe Nagorye is inhabited by
bright butterflies almost twice as large [as E. iduna sajana
Higgins, 1950], the females being larger than males.
Specimens occur among them with a darker pattern, alike ab.
sulitelmica Schultz, 1906. These large butterflies are
designated here as E. iduna semenovi Korshunov et Ivonin,
sbsp.n.
HOLOTYPE: a female. F.w.l. 24 mm. The pattern is as in the
main form, but light elements are more distinct and
suffusion with dark scales is diminished on them. There are
small dark spots at the vein ends.
ALLOTYPE: a male. F.w.l. 20 mm. The light elements of the
pattern above are suffused with grey and dark scales,
especially at the fulvous band. Beneath the light elements
are as in the female.
MATERIALS: The holotype: a female - 14.7.1975, Khakassia,
surroundings of the mountain Bobrovaya, a stream being a
tributary of the Sarala River (Yu. Korshunov). The allotype:
a male - 7.07.1975, the same locality, on a dandelion
inflorescence. Paratypes: 47 males 15 females - 7-
14.07.1975, the same locality; 5 males 3 females, 30.06-
9.07.1996, the Kedrovaya River Valley at the mountain
Chemodan; 8 males 1 female, 4.07.1996, the valley of the
Krasnaya Rechka River, a tributary of the Srednyaya Ters'
River; 5 males 3 females - 5.07.1996, the mountain Stanovoi
Khrebet [not to confuse with homonymic mountain range in
Transbaikalia]; 1 male - 6.07.1996, a lake at the mountain
Medvezhya; 1 male 1 female - 8 and 10.07.196, meadows at the
mountains Chemodan and Lysaya (V. Ivonin).
In general, the butterflies prefer meadows with Viola
altaica on mountains slopes and at screes. They were noticed
to feed on the flowers of valeriana, Bistorta bistorta,
Ranunculus, taraxacum, rarely on Umbelliferae.
The subspecies is dedicated to the memory of our teacher
Semenov, Boris Sergeevich, which as long ago as in 20s
collected Lepidoptera in these places and discovered
Argynnis sagana in the ters' Rivers basin. For a long tome
he was a head of the Nature division of the Novosibirsk
Museum of Local Lore. "
Euphydryas iduna alferakyi Korshunov [et P. Gorbunov,
that was omited in the publication - Yu.P.] 1996
Original description:
"...Yakutian butterflies differ from others, They look
mottled. differ from the typical ones by pure white and
brick-red spots on both wing sides and a larger size. We
name them Euphydryas iduna alferakyi Korshunov, sbsp.n.
HOLOTYPE: a male. F.w.l. 19 mm. There is a transversal
band of black scales on the white field on the wing
upperside. The wing undesride have a contrasted pattern,
characteristic to the species, and black veins.
ALLOTYPE: a female. F.w.l. 23 mm. Close to the male in
the pattern and colouration but has more black scales on
the white areas on the wing upperside.
MATERIALS: The holotype: a male - 21.06.1985, E Yakutia,
18 km NEE of the settlement Khandyga, the Khandyga River
headwaters (V. Dubatolov). The allotype: a female,
12.06.1985, E. Yakutia, the ALdan River, Khandyga, a damp
clearing in a larch forest (V. Dubatolov). Paratypes: 2
males, 26.06.1971, the settlement Argatakh on the
Srednekolymskii District of Yakutia (P. Polyakova); 2 males
1 female, 16.06 - 15.07.1973 and 1 female - 20.06.1979, the
In'yali River headwaters, Yakutia (N. N. Vinokurov); 4 males
- 12.06.1985, the Aldan River; 11 males 5 females - 21.06-
17.07.1985, the Khandyga River headwaters; 7 males 2 females
- 5-20.07.1985, the Suntar River low reaches, the Indigirka
River basin; 8 males 3 females - 17-18.07.1985, the Kyubyume
River, the Indigirka basin (V. Dubatolov); 1 male -
30.06.1985 - the Suntar-Khayata mountain range, the mountain
baranya (L. Popova); 1 male - 15.07.1990, E yakutia, the
Myurale River headwaters (M. Zakharov).
The subspecies is named in the honour of Sergey
Nokolaevich Alferaky (1850-1918), an author of works on
Siberia and a great number of taxa."
Melitaea menetriesi westsibirica Dubatolov in Korshunov, 1998.
Original description:
"GEOGRAPHICAL VARIATION. Subspeceis westsibiricva Dubatolov, sbsp.n. -
Verkhnee Priobye [the Upper Ob' basin]. F.w.l. 14-19 mm. Externally little
differs from the subspeceis from the South Siberiam mountains, resembles
also aurelia. The main feature - tegumen in aurelia without distinct teeth,
in westsibirica the teeth are long while aedeagus apically has a transparent
prominence bearing a curved upwards processus, as in aurelia. These butterflies
occur sympatrically at Karasuk and southwards, in the Altaiskii Krai Province.
MATERIALS: Holotype - a male, 29.06.1957, the village Novyi Sharap of
Ordynskii District of Novosibirsk Province (Y. Korshunov). Paratypes - 3
males, 22.06.1958, 25.06.1959, the same label, a female,
17.06.1982, village Acha of Bolotninskii District of Novosibirsk Province
(Y. Korshunov), a male, 8.06.1994, village Nizhnii Koen of Iskitim
District of Novosibirsk Province (V. Ivonin)".
Melitaea latonigena ukoka Korshunov, 1998.
Original description:
"Below data are given about an isolated population of latonigena on
the Ukok Plateau (SE Altai) in the Kalguty River basin. In extremal conditions
of highland biotopes among bunch-grass tundras on a height above 2000 m.
A. V. Bondarenko found out more than two dozens of specimens which we
name here as
M. latonigena ukoka Korshunov, sbsp. n.
Males differ by a partai reduction of black pattern, females by a distinct
black chequered pattern on the light ground colour.
HOLOTYPE. A male of 11.07.1997, f.w.l. 17 mm, of black spots on fore wing
well develped are marginal ones - at light fringe and next, and then only
in the middle of the wing close to the base. So the outer field lacks
the spots completely, while in all other forms of latonigena this
area of the wing has either black dots between the veins or even dots
fused into a chain, as in atrata. In one male of the paratypes the
central row of spots is also not expressed.
ALLOTYPE. Female of 11.07.1997. F.w.l. 18 mm, on a chequered pattern of fore
wing a light spot in cell, situated basally of discal vein, is
distinct. It is quadrangular, in some paratypes of different shape but
invariably well expressed. In the allotype on hind wing upperside
several plates situated closer to fore margin are orange coloured,
in paratypes some parts of fore wing also have the same colour. Pattern
of wing underside is of a latonigena type. Among specimens of the Ukok
there are several aberrant females in which wing upperside is entirely or
partly black and on wing underside with a black pattern developed in basal area.
In total the type series embraces three males and seven females, with f.w.l.
16-20 mm. Besides, I enumerate similar butterflies from the collecctions of
R. Yakovlev and V. Doroshkin, also from the Ukok Plateau. 5 females, 5 males
- 14.07.1994, 1.07 and 5.07.1995, 25.06.1996, 10 and 15 km S of village
Szhazator, 2200 m; 4 female and 35 males - 3.07.1997, mountain Mai-Tobe, 2600
m; 2 females and 18 males - 10-12.07.1997, middle flow of the Dzhumaly
River, 2200 m."
Meltaea sutschana graeseri P. Gorbunov, 1995.
Original description:
"...Small (f.w.l. 18-20 mm) and very peculiar butterflies
from the Sakhalin Island are described as Meltaea sutschana
graeseri P. Gorbunov, sbsp.n.
HOLOTYPE. A male. F.w.l. 20 mm. The wing upperside ground
colour is ochre-orange, more faded than in the typical
butterfliws. Among the black marking a row of spots on the
middle of the wings, these spots are isolated from each other
and more lengthwise elongate than in sutchana. On the fore wing
upperside there are only four (also lengthwise elongate) spots
at the fore margin. The hind wing underside ground colour is
white with a yellow tint; the ochre-orange bands are wide,
occupying about a half of the wing area. The light space between
the external and the internal bands does not exceeds the width
of the former (about 3 mm). In this space the black spots are
close to the internal band, while in sutchana they are closer to
the external one. The antennae and their clubs are fulvous. The
genitalia structure is as in M. sutchana.
MATERIALS: The holotype: a male - 2.07.1994, the Sakhalin, the
environs of the city Yuzhnosakhalinsk (A.M. Basarukin). The
paratype: a male - 2.07.1994, the same locality.
We include only two specimens from the Sakhalin into the type
series, although a male of a similar appearance was found also
in our collections from the Lower Amurland (the Komsomol'skii
Reserve, Bichi, 23.06.1988), and two more such males - in the
ISEA collection from the southern Transbaikalia (Sokhondinskii
Reserve, the Agutsa River middle flow, 9.08.1987). These facts
rise a supposition of a species rank of the taxon being
described, that need further materials for corroboration."
It should be added that the subspecies is named in the honour
of Ludwig Karl Fridrich Graeser (1840-1913), a German
lepidopterologist who studied butterflies on the Amur River in 1881-
1885.
Melitaea fascelis singularia Korshunov, 1995
The name was proposed in Korshunov & Gorbunov (1995) as
follows: "...Yu. P. Korshunov (1977) described a peculiar female
of M. fascelis from the surroundings of Lake Hubsugul in
Mongolia. We suggest the name Melitaea trivia singularia
Korshunov, sbsp.n. for such butterflies." There was an obvious
error in the species name, which should be fascelis instead of
trivia, as it is clear from the preceding words as well as from
the species headline: "Melitaea fascelis (Esper, 1784) (= trivia
Denis et Schiffermuller, 1775)" Translation of the cited
description, which should be considered as the original
description of the subspecies, is follwoing:
"*Melitaea trivia Denis et Schiffermuller, 1775, ?sbsp.n.
Hubs[ugul Aimak]: Lake Hubsugul, 26 VI (D. [V.Ch.
Dorogostaiskii], 1 female.
From the typical form differs by light and mottled pattern,
somewhat elongate wings. Generally the habitus resemble that of
M. t. ignasiti Sagarra, 1926. Probably, belong to a new
subspecies.
Female. The palpi are set with yellow hairs. The fore wing
lengthis 18 mm. The wing upperside is pale-fulvous, mottled
since black dots are small and well contrasted. On the hind wing
underside the black strokes are narrow, do not contact
completely to the fulvous band. The middle field is light-
yellow, not differing in colour from the basal and outer parts.
There are two rows of four irregular black spots (dots) going from
the fore margin to the cell in the middle field. The genitalia
are close to those in the typical form."
Melitaea uvarovi P. Gorbunov, 1995.
Original description:
"173. Melitaea uvarovi P. Gorbunov, sp.n. [Shashechnitsa
Uvarova]
The species is being stated by five males collected by S.V.
Shutov in the valley of the Ural River, together with such
spring steppen species as Zerynthhia polyxena, Triphysa phryne,
Proterebia phygea [an error, should be "Proterebia afra" - Yu.
P.], Thersamonia thersamon etc. From a close species M.
fascelis, the closest population of which is known in 15 km NE
of the place of collection of the new species (12 km S of the
station Kuvandyk) it differs by a great number of external
characters, and also early flight period (over-wintering in a
phase of pupa is probable). It is not excluded that the taxon
deing described will turn out to be conspecific to a Central
Asian species M. robertsi Butler, having a certain similarity
with it in appearance and life history. But now, taking into
account the Ural populations to be so remote, we describe it in
a species rank.
MALE: F.w.l. 15.2-16.4 mm (15.4 mm in the holotype). The
wings are somewhat narrower than in M. fascelis. The fore wing
ground colourt is ochre-orange-red, noticeably darker than in a
close Central Asian taxon M. robertsi catapelia Staudinger,
1886. The black spots on the wing upperside are smaller and more
clear-cut than in M. fascelis. The black marginal band is about
1 mm wide. A row of black submarginal lunules goes along it on
both wings, they may contact to the marginal band but do not
form, together with it, an united marginal blackeinig, found in
M. f. fascelis. There is a small white spot on the fore wing
fore margin (3 mm off the apex). The basal blackeinig on the
hind wing does not extend above the vein Cu1. The fore wing
underside is ochre-orange with a whitish spot at the apex and
rather small isolated black spots. The hind wing underside
ground colour is white, without a yellowish tint peculiar to M.
fascelis. The pattern is ain in M. fascelis but the black spots
at the outer margin are narrower. The fringe is white, more
clear and long than in M. fascelis, with black patches at the
vein ends. In the genitalia structure the differences from M.
fascelis (see Appendix) are rather weak.
MATERIALS: The holotype: a male - 6.05.1982, the Orenburg
Region, the village Donskoe, a valley steppefied meadow (S.V.
Shutov). Paratypes: 4 males - 3-6.05.1982, the same locality.
The species is dedicated to the memory of Uvarov, Boris
Petrovich (1888-1920), a honoured member of the All-Union
Entomologiocal Society, who studied the fauna of Lepidoptera of
NW Kazakhstan from a young age."
Clossiana chariclea tshuktsha Dubatolov Dubatolov et Korshunov, 1998.
Original description:
"In Chukotka and Kamchatka there is ssp. tshuktsha Dubatolov et
Korshunov, sbsp. n. In general it resembles butleri by colouration
but well differs by the male genitalia structure. In the new subspecies
the outer area of hind wing underside is evenly red-brown, in males
a yellow colour presents only in some specimens, between veins M2 and CuA
outside of dark postdiscal spots. The fore apical processus of the valva
is shortened, no longer than the lower one (in butleri this processus
is very short and more stout).
MATERIAL. Holotype - a male, 2?.07.1986, 100 km S of Pevek, E Chukotka,
Ust'-Chaun (Dubatolov, ZInchenko). Paratypes - 12 males, 14 females, the
same locality, 25.07-2.08.1986 (Dubatolov, Zinchenko). The fact that specimens
of a new subspecies in general have the genitalia intermediate between
chariclea and butleri gives a reason to consider butleri W. H. Edwards, 1883
as a subspecies of a circumpolar chariclea."
Clossiana astarte ershovi Korshunov et P. Gorbunov, 1995
Original description:
"...Butterflies from the Prilenskoe [near the Lena River]
Plateau differ by relatively large size (f.w.l. is 22-24 mm in
males, 23-26 mm in females) and the pattern of the hind wing
underside.
Clossiana astarte ershovi Korshunov et P. Gorbunov, sbsp.n.
HOLOTYPE: A male. F.w.l. 22 mm. The wing upperside is fulvous
with an intensive black pattern; the hind wing is substantially
blackened in the basal half. The wing underside is mottled, a
row of round black spots at the outer margin and their light
rims are especially conspicuous. On the hind wing underside the
outer half ground colour is reddish-brown; the discal band on
the hind wing is more mottled than in other subspecies, it
contain contrasted patches of whitish and brown colour.
MATERIALS: The Holotype : a male - 19.06.1968, the settlement
Khaptagai on the Lena River (Yu. Ammosov). Paratypes: 2 males -
23.06 and 11.071987, the settlement Khaptagai, the road to Tammu
(Yu. Ammosov); 2 females - 20.07.1962, Yakutsk, the botanical
garden (Alexandrova); a male - 7.06.1979, the same locality; a
ma;e - 8.06.1985, the same locality (V. Dubatolov), a female -
23.06.1992, the same locality (P. Gorbunov).
The subspecies is dedicated to the memory of Ershov,
Nikolai Grigoryevich (1837-1896), one of the first real members
of the Russian Entomological Society, the author of numerous
works and the first catalogue of the Siberian [an error - must be
Russian - Yu.K.] butterflies."
Clossiana astarte suntara Dubatolov, 1998
Original description:
"V. V. Dubatolov finds it necessary to isolate a subspecies the butterflies
of which keep to rocky slopes and screes of the Suntar-Khayata mountain
range. This is suntara, Dubatolov, sbsp. n. F. w. l.: 17-27 mm.
The hind wing above the fulvous ground colour almost excluded with
black scales, so that a par of large postdiscal spots remain surrounded
with narrow or wide light rims. Dark rimming of medial band slightly
darker than fore wing underside ground colour.
MATERIALS. Holotype - a male, 07. 1985, E Yakutia, the Suntar-Khayata
mountain range, 180 km ENE of the highway Khandyga-Magadan (Dubatolov,
Barkalov, Popova)."
Clossiana matveevi P. Gorbunov et Korshunov, 1995.
Original description:
"MALE: 22.6-23.8 (in holotype 22.8 mm). The wing upperside
is ochre-orange, more faded than in C. tritonia and C. erda and
close to that in C. astarte. The black pattern in general
corresponds to that in C. tritonia but is noriceably fine; a
more narrw black margin without widening at the veins, as is
characteristic for C. tritonia, is among significant
differences. On the fore wing underside the pattern is the same
as on the upperside, but the black spots are more narrow and two
black parallel lines replace the margin. The hind wing underside
ground colour varies from muddy-orange to muddy brick-red; the
ground-ochre or greyish discoidal band is relatively narrow,
2.2-2.5 mm wide at the vein M1 (in other taxa of the tritonia-
group it is not less wide than 2.5 mm), its both sides being
bor dered with conspicuous black streaks, and locally it is much
suffused by dark scales, which entirely hide it at the anal
margin. The postdiscoidal area contains a row of seven well-
expressed whitish spots. Outside of it there is a row of six
black spots and then the area of ochre-red colour 2.0-2.5 mm
wide. The black bracked-shaped submarginal spots are distally
margined with white spots streched out across the veins; outside
of them there is a double marginal line. In the male genitalia
the caudal processus of the valva ends with a single large
tooth, not with a row of small teeth as in C. astarte and C.
tritonia, the costal processus is large, foot-shaped (for the
figure see Appendix).
MATERIALS: the holotype: a male, Altai, the Kurayskiy mountain
range. the settlement Aktash, 2700 m above sea level, a dry
scree, 8th July 1990 (E.A. [must be E.M. - O.K.] Matveev leg.).
Paratypes: two males one female, the same locality 16th July 1992;
4 males - the Kuraiskiy mountain range, the Yarly-Ayry river
headwaers, 3000 m altitude, 7-12th July 1992 (Yu. Prokofyev).
The taxon is named by the surname of Evgenii Aleksandrovich
[an error - must be EMvgenii Mikhailovich] Matveev, a Moscow
naturalist, a butterfly lover, who first collected these
butterflies. Although it is not excluded that a butterfly of this
very species (identified by A.A. Meingard as C. amphilochus
Menetries) was caught by V.V. Sapozhnikov on 6th June 1905 in the
Tsagan-Kol River headwaters in Mongolia."
Clossiana eunomia riamia Korshunov et Ivonin, 1998.
Original description:
"Peculiar turned out populations of the ryams [loc. raised peat-moss bogs within
the forest-steppe zone] of the Novosibirsk Province. Imagines keep there to
Eriophorum and Ledum, often resting on their leaves and pine needles, they
are active both in sunny and overcast weather. the butterflies of ryams are
large, f. w. l. not less than 18 mm, more frequently 20 mm. The brick-red
individuals are rather melanized, the spots of black pattern enlarged,
wing margins black-rimmed on which blueish spots are scarcely seen. On hind
wing underside ocelli and light spots bright silvery-blueish-white, in many
specimens of the same colour are the spots of marginal row on hind wing
underside. Note that such butterflies were met with still by the expedition
by N. F. Kashchenko on 8th June 1899 in a ryam at the station Ubinskaya where
more than 30 specimens were captured. A. A. Meinhard (1905) in his "List VI"
did not give them a special name bot noted a dark ground colour
and large and fusing with each other black spots and bands on the upperside
of all the wings and a very bright reddish-brown colour of the hind
wing underside with conspicuous large spots with a strong silvery-nacreous
glittering. Among 30 specimens two had "unusually dark colouration" and
five more light, closer to the typical eunomia.
The new subspecies is name riamia Korshunov et Ivonin, sbsp. n.:
MATERIALS: Holotype - a male, 17.06.1997, Kamennyi Ryam at v. Kuznetskoe
of the Chulymskii District of Novosibirsk Province (V. Ivonin), allotype -
a frmale, 17.06.1997, the same locality. Paratypes - two males, 17.06.1962,
a ryam at v. Kaily of the former Mikhailovskii (now Ubinskii) District
(Y. Korshunov), one female, seven males - 17.06.1997, Kamennyi Ryam,
26.06.1997, the same locality."
Clossiana eunomia stromi Korshunov, 1998.
Original description:
"It turned out that in the mountains of S Siberia there occur two subspecies.
For Altai, the Sayans, Tuva and adjacent parts of Mongolia a new subspecies
stromi Korshunov, sbsp. n. is described (by the name of Semen Aronovich
Strom (1906-1985), my main expedition teacher, a head of the youth geographical
society "Sibir'" in Novosibirsk). As different from acidalia the butterflies
of this subspecies lack the silvery-white colour on the wing underside.
From the nominotypical one they differ by a smaller size (f.w.l. usually
less than 20 mm), they have a finer black pattern, especially in males.
Lastly, for stromi, characteristic is a presence of a large ochre-red
spot (or also a small one near) at basal darkening of the hind wing upperside.
On the hind wing underside the colour of rather small ocelli does not differ
from that of all other light spots (light salad-coloured or slightly yellwish).
These light spots are surrounded with ochre .... [the word was missing in
the dersciption].
Material: Holotype - a male, 24.07.1975, W Sayan, Pazyryk-Ergak-Taiga
mt. tange, 1st brook behind the pass [from which side remains unclear]
(Y. Korshunov), allotype - a female, the same label. Paratypes - a
male, 23.07.1948, Tuva, W Tannu-Ola (A. I. Cherepanov), a female.
9.07.1954, Altai, v. Abai, two males, 14.07.1959, Chuiskaya Step',
the Yustyd River at v. Ak-Tal, two males and two females,
19.07.1963, by the village to Ust'-Ulagan (O. Ivanovskaya), a male,
6.07.1966, the Mezhtyyaryk River - a tributary of the Chuya, 1700
m (A. E. Standel), a male, 7.07.1966, Altai, mountain Sukor at
CHagan-Uzun, 2600 m (A. E. Shtandel), a male, 70 km SE of Lake
Dood-Nur, 2000 m, Mongolia (V. Solyanikov), a female, 21.07.1972,
Tuva, lake Kara-Khol', a male, 5.07.1976, Altai, Seminskii Pass,
two males, 16.07.1982, Altai, a male, 11.07.1983, Katunskii mt. range,
stow Bertkhem, 15 km SW of Katanda (V. Dubatolov), a male and female,
13.07.1983, 15 km SW of Katanda, a pass between the Kuragan
and Kucherla Rivers, Tukura, 2400 m (V. Dubatolov), a male and two
females, 20-21.07.1983, Terektinskii mt. range, 10 km N of Katanda,
highlands, 2500 m (V. Dubatolov), a male, 12.07.1987, Lake Teletskoe,
mountain Kolyshtu (A. Barkalov), a male, 16.06.1987, Tuva, Todzha,
Lake Azas (V. Zinchenko), two males, 25-29.06.1991, Sayano-Shushenskii
Reservation, the Ak-Sug River headwaters and the Ala-Aya River middle
flow (V. A. Vagin). Further in the mountains of Siberia, Yakutia and
in the northern Far East there ranges te subspecies acidalia Bober, 1809
(=asiatica Staudinger, 1901) described from Pribaikalye, which differs
from the nominotypical one by a smaller size, a black pattern closer
to ossiana, as well as the underside with expressed silvery-white
spots. AS to the northern subspecies ossiana Herbst, 1800 proper,
it embraces small butterflies with the ocelli always differing in
colour from the background (light or with a silvery tint) and extends
at least to the latitudes of the Taimyr."
Boloria altaica pustagi Korshunov et Ivonin, 1995
Original description:
"By the materials from Gornaya Shoria [elevation] the
subspecies Boloria altaica pustagi Korshunov et Ivonin, sbsp.n.
is being described, differing from altaica first of all by a
darker (brick-red) ground colour.
HOLOTYPE: a male. F.w.l. 21 mm. The wing upperside is brick-
red with the black pattern enlarged, as compared with the
subspecies altaica. The black spots along the outer margin are
not fused into a streak. On the hind wing underside the discal
band is bright, well contrasted to the ground brick-red
background; the black round spots in the postdiscal area are
contrasted, some of them contain white nucleoli.
In females the upperside is lighter, yellowish, the underside
is bright, as in males.
MATERIAL: The holotype: a male - 13.07.1992, Gornaya Shoria,
the environs of the settlement Sheregesh, the mountain Pustag,
1000-1300 m (V. Ivonin). Paratypes: 7 males, 3 females -
13.07.1992, the same locality."
Triphysa albovenosa sacha Korshunov, 1996
Original description:
"HOLOTYPE: a male. F.w.l. 18 mm. The wing upperside is
blackish-grey with a steel flash, without any pattern, only the
dot or ocelli of the underside are slightly seen through. The wing
underside is as in T. albovenosa, but there are small dots or
ocelli in the outer field, the second one of which on the hind
wing even has a white pupil. There are three dots and one
ocellus on the fore wing, the hind wing has five small black
ocelli between the veins, the third from above being a dot. In
some paratypes the dots are only on the hind wing, in two
paratypes they are hardly noticeable at all.
ALLOTYPE: a female. F.w.l.: 15 mm. The wing upperside is
greyish-white, the underside pattern is seen through. On the
wing underside there are very small black dots: four on the fore
wing and five on the hind wing. The wing area between the outer
field and the base is mottled due to grey spots of different
size. In other females these grey spots fuse to each other, in
one female the wing underside is as in the males, with the dots
large, especially on the fore wing.
MATERIALS: The holotype: a male - 7.06.1985, Yakutsk, the
botanical garden (Dubatolov). The allotype: a female -
9.06.1985, the same locality. Paratypes: females and 2 males -
13.06.1968, Yakutsk, Chugur-Murun; a male and a female -
10.06.1974, Khaptagai, the Tamma River valley (Ammosov); a male
- 1.06.1977, at the settlement Oktemtsy; 3 males 1 female -
28-31.05.1989, Yakutsk, the botanical garden (Dubatolov)' 10
males 10 females - 7-9.06.1985, the same locality; a male -
1.06.1985, 5 km of Yakutsk on the road to Magadan, a bogged
rivulet valley (Dubatolov).
So, the range of a new subspecies lies within Yakutia,
noteworthy that the Suntar Khayata mountain range in E Yakutia
is inhabited by different butterflies, close to the subspecies
tscherskii Crum-Grshimailo, 1899. It has a somewhat darker
ground colour on the wing underside, the light veins being more
conspiculus as compared with other subspecies. This subspecies
ranges widely in E. Siberia, it was also found in Transuralia
and West Chukotka."
Coenonympha amaryllis borisovi Korshunov et Ivonin, 1996.
Original description:
"...The largest and peculiar butterflies among amaryllis have
been found in Pribaikalye, which are secribed as C. amaryllis
borisovi Korshunov et Ivonin, sbsp.n.
HOLOTYPE: a male. F.w.l.: 20 mm. The wing upperside is ochre-
yellow. There are two large and two small black dots on the fore
wing and five large dots and a dark marginal rim on the hind
wing. On the underside there are very large, contrasted ocelli
with silvery pupils and a relatively wide silvery streak along
the margins. The fringe is light. On the most part of the
species the butterflies have the underside without any pattern,
that on the underside is more dull, the ocelli are smaller than
in borisovi.
MATERIALS: The holotype: a male - 24.06.1988, Baikal, the
village Listvyanka (Ivonin). Paratypes: 2 males - 30.06.1983,
the same locality (Yu. Timoshenko), 3 males -24.06.1988, the
same locality - 20.06.1991, the Verkhne-Angarskii mountain
range, the environs of Severobaikal'sk (Ivonin). All the
butterflies were collected on steppefied mountain slopes.
The subspecies is named by the surname of a Decemberist
Borisov, Petr Ivanovich (1800-1854), the author of first aquarel
portraits of Lepidoptera of Baikal."
Hyponephele cadusina gurkini Korshunov, 1995.
Original description:
"... A new subspecies Hyponephele cadusina gurkini Korshunov,
sbsp. n. is described on the materials from Central Altai,
differing from the nominotypical one by the following
characters: on the fore wing upperside the yellow field is pale,
the dot between the veins Cu1 and Cu2 is larger, the dark margin
on the hind wing underside is wider, the wing underside is more
evenly coloured, the dots at the anal angle are smaller.
HOLOTYPE: a male. F.w.l.: 20 mm. The wing upperside is
brown, the hind wing is evenly coloured, the fore wing has a
wide ochre-yellow spot on the outer field. In the apical part of
this spot there is a black ocellus with a light pupil and a
twice as small black dot below it, between the veins Cu1 and
Cu2. A black androconial brand is covered by brown scales along
its fore and hind margins. The fore wing underside is ground-
yellow with a relatively wide dark marginal band and a light-
rimmed apical ocellus. The hind wing underside is grey with
brown marble which does not form transversal bands, as is
observed in cadusina.
MATERIALS: the holtype - 1st July 1974, Central Altai, a
steppefied bank slope of the Katun' River near its junction with
the Chuya River, (Yu.P. Korshunov leg.). Paratypes: - 2 males,
1st July 1974, the same locality.
The subspecies is dedicated to the memory of an Altai
painter Gurkin, Grigorii Ivanovich 91869-1937)"
Erebia euryale flaveoides Korshunov et Tatarinov, 1996.
Original description:
"...In the lowland taiga of the north-east of Europe the
subspecies euryaloides Tengstrom, 1869 ranges (described within
ligea), in which both sexes have white bands. A form arctica
Poppius, 1906, also with a white band, is known from the Kanin
Peninsula, later the taxon boreomontanum Sedykh, 1977 was
described from the Sob' River headwaters, which is a synonym to
arctica. The North< Subpolar and Polar Ural and the
Bol'shezemel'skaya Tundra are inhabited by the butterflies of
which the females have bright brownish-yellow band. These
butterflies are described here as the subspecies flaveoides
Korshunov et Tatarinov, sbsp.n.
HOLOTYPE: a female. F.w.l. 21 m. On the hind wing underside
tere are two small brown spots on a brownish-yellow band: the
upper at the incision in the band at the cell apex, the lower
between the bein Cu branches.
ALLOTYPE: a male. F.w.l. 20 mm. On the hind wing the band is
reduced to a white spot, situated where a female would have
an incision of the band, and some white scales above and below it.
MATERIALS: The holotype: a female - 15.07.1995. The Subpolar
Ural, the middle flow of the Malyi Patok River [(Tatarinov) - was
omited Yu.K.]. The allotype: a male - 16.07[.1995], the same locality.
Peretypes: 9 males 9 females - 15-17.07[.1995]. Only one point on
the Polar Ural is known, the station Krasnyi Kamen', where
butterflies of both types [with white and brownish-yellow band
in females] occur together, but this is at a railway, which was
a channel of penetration there the butterflies from euryaloides
populations."
Erebia kosterini P.Gorbunov, Korshunov, Dubatolov, 1995.
Original description:
"253. Erebia kosterini P.Gorbunov, Korshunov, Dubatolov, sp. n.
[Chernushka Kosterina].
TYPE LOCALITY: The Khindzha River on the Koni Peninsula (the
Magadan Region)
BIOTOPES: valley meadow plots between the thickets of the bush
alder and the dwarf pine.
FLIGHT PERIOD OF IMAGINES: observed in the second half of July.
HOLOTYPE: a male: F.w.l.: 21.5 mm. The wing upperside is
brown with a wide (3-4 mm) ochre-fulvous band in the postdiscal
area, which contains: a black dot at the apex between the veins
R5 and M1, a double black spot between M1 and M3, the next [from
the apex] intervein plate contains a dot, the next one contains
a round black spot. No adroconial brand is seen. On the hind
wing upperside there are four round black spots on an ochre-
fulvous band going from the fore margin to the vein Cu2. The
third spot (counting from the fore margin) is noticeably shifted
outside. The underside is analogously coloured, but the ground
colour is darker to some extent, the band on the fore wing is
ochre-yellow, while the black dot at the apex is absent; on the
hind wing the spots are smaller and reside not on the band but
in narrow ochre rings, inside of which there is a white band
split into fragments. The fringe is chequered. In the genital
apparatus the valva is narrow (the ratio of its maximal width to
the length being 1:5), slightly and gradually tapering to the
apex; a half of its dorsal edge bears teeth of various size (see
Appendix). In a similar species E. ligea the valva is sharply
(two- or three-fold) narrowed at the apex forming a finger-like
projection. A structure of the male genitalia similar with that
of E. kosterini is found in an American species E. vilderi
Elwes, 1898 [an error in the original description, must be
vidleri - Yu. K], well differing by the absence of postdiscal
black ocelli on the hind wings.
ALLOTYPE: a female. F.w.l.: 20 mm. The wings are noticeably
narrower than in the holotype. The colouration and pattern are
similar to those of the holotype, differing in the following: on
the fore wing upperside the black dot at the apex is absent, on
the hind wing upperside the black spots are smaller and there is
only three of them (that colsest to the fore margin is wanting);
the double spot on the fore wing upperside, the same double spot
and the spot between Cu1 and Cu2 [on the fore wing underside -
these words were accidentally dropped in the printed version -
Yu. K.] , and three of the four spots on the hind wing underside
have white nuclei. On the hind wing underside the white stripe
inside of the row of ocelli is brighter and more clear-cut than
in the holotype.
MATERIALS: Holotype: a male - 20.07.1989, the Magadan Region,
the Koni Peninsula, the lower reaches of the Khindzha River, a
herbaceous meadow on the lower terrace (O.E.Kosterin). Allotype:
a female - 20-24.07.1989, the same locality. Paratypes: 2 males
2 females - 20.07.1989, the same locality and date.
The species is named by the surname of Oleg Engel'sovich
Kosterin, a participant on numerous expeditions in Siberia and
the Far East, who collected the type series of butterflies of
the species."
Note: a more detalied description of the habitat of these
butterflies (referred to as Erebia (ligea (Linnaeus, 1758))
group) on the Koni Peninsula can be found in (Kosterin, 1993).
After the first description the butterflies of this species were
collected also at the Magadan City and the settlement Karamken in
the Magadan Region (Korshunov, 1996). Later, it turned out that
this name is a junior synonym of Erebia ajanensis Menetires: see
(Dubatolov et al., 1999)
Erebia rossi subarctica Korshunov, 1996.
Original description:
"...The butterflies from the zonal tundras and forest-tundras of
N Asia and from mountains of Yakutia and Chukotka are different.
They differ first of all by the absence of a reddish-castane-
brown colour on the fore wing underside in a majority of
specimens. Only some females have a brand of this colour and few
males have groups of such scales at the ocelli. We describe them
as Erebia rossi subarctica Korshunov, sbsp. n.
HOLOTYPE: a male. F.w.l. 24 mm. The wing upperside is brown,
there is a double fulvous spots at the fore wing apex,
containing two black dots, and a a hardly noticeable fulvous
spot without a dot at the Cu veins. The hind wing upperside has
no dots. On the fore wing underside there is a double fulvous
spot with two black dots centered with white ocelli, a tiny
fulvous speck and a small fulvous spot, with a white-centered
black dot, between the branches of the vein Cu. On the hind wing
undeside there is an obscure band, with a white dot on its marhin
at the cell apex, and a somewhat ligher band in the outer field.
There is a "flash" of reddish scales at the spots, noticeable
only if to place the butterfly at different angles to the light.
ALLOTYPE: a female. F.w.l. 25 mm. The wing upperside is as
in males, but the small spot at the Cu is missing. The fore wing
upperside has only two black dots with white pupils in a common
fulvous round spot at the apex. The hind wing underside has the
same bands as the male, the outer margin of the outer one
bearing two very small white spots.
Among the paratypes about a half of specimens has 2-3 spots
in addition to the double apical one on the fore wing and 2-4
Spots on the hind wing. On the hind wing underside there may be
fore white dots. The imagines from the Maymecha River basin and
from a more northern Novaya River, on the Putorana Plateau, are
peculiar: they are smaller, f.w.l. being 19-21 mm, and have
only the apical spots (the only female of 8 specimens has spots
also on the hind wing upperside), this population deserves
further studies.
MATERIALS: the holotype: a male - 15.07.1968, the left bank
of the Tanlovaya River, a tributary of the Schuchya River, S
yamal (Korshunov). The allotype: a female - 14.07.1968, a tundra
between the Tarseda River and the Schuchya River left bank, S
Yamal, Korshunov). Paratypes: 10 males 7 females - 14-
30.07.1968, the Schuchya River basin (Yu. Korshunov); 2 males -
28-29.06.1969, the Omolon River valley (N. Gomoyunova); 2 males
- 8.07.1976, the Yenisei River lower reaches, Chopko
(Chernenko); 2 males - 07.1981, Taymyr, Ary-Mas, Novaya (R.
Polovinkina); 1 male 5 females - 28.07-8.08.1982, Putoran,
Maymecha (I.S. Zakharzhevsky); 1 male - 22.06.1984, the Sutar-
Khayata mountain range (L. Popova); 14 males 1 female - 19.06-
14.07.1985, the Suntar-Khayata mountain range, the Khandyga
River headwaters (V. Dubatolov); 1 male - 18.06.1989, Yakutia,
the Kele River headwaters, a Carex/Eriophorum tussocks (L.
Popova); 2 males - 12-17.07.1993, the Chita Region,, 35 km SSE
of Chara, Naminga (A. Streltsov)."
Erebia mackinleyensis ola Korshunov, 1995
Original description:
"... Specimens from the Magadan Region are characterized by
a complete vanishing of castane-coloured spots on the wing
upperside in males and their reduction in females. Below we
describe them as a new subspecies. The shape of the valva in the
male genitalia in sachaensis and ola is very variable, that is
illustrated by the drawings of the genitalia of the holotypes
(see Appendix). In general, the genitalia of these taxa have no
significant differences between each other, as well as from E.
semo and E. erynnin.
Erebia mackinleyensis ola Korshunov, sbsp. n.
HOLOTYPE: a male. F.w.l.: 25 mm. The wing upperside is evenly
soot-coloured. The underside is of the same colour but on the
fore wing, at the cell, there are groups of reddish scales
forming a hardly detectable sinking spot. The light scales are
scattered over the hind wing underside where no bands are seen.
In the genitalia the valva has a rounded apex.
ALLOTYPE: a female. F.w.l.: 26 mm. The wing upperside is black
with a brown tint. On the fore wing, at the cell, there are
reddish elongate strokes between the veins. On the fore wing
underside the corresponding reddish spots are bordered with dark
scales. The hind wing underside is not only mottled, as in
males, but have also distinct traces of the postdiscal band, the
margin of which is most clearly seen in the middle part of the
outer field.
MATERIALS: the holotype: a male -7.07.1991, the Magadan
Region, the headwaters of the Ola River, the surroundings of the
mountain Nukh (1828 m) (V.V.Palekha leg.); the allotype: a
female - 7.07.1991, the same locality; paratypes: a male -
07.1991, the same locality; 2 males 1 female - 13.07.1993, the
same locality; a male - 6.07.1964, the Omsukchanskiy mountain
range, at the setlement Golimyy, 6th July 1964 (A.V.Tsvetaev)."
The attribution of this taxon to E. mackinleyensis as a
subcpecies, made by P. Gorbunov in a published version of the
book (Korshunov, Gorbunov, 1995), is erroneous. I consider it to
be a bona species due to the complex of characters concerning the
general appearance, pattern, the details of the genitalia
structure (Korshunov, 1996). New species has a certain similarity
to E. erynnin Staudinger, 1894, but this species of the E Sayan
lacks androconia resembling those of E. ola, their genitalia
structure have substantial differences, in particular, the end of
the valva is twice as thick in E. ola, in general the valva is
thicker and resembles that in E. mackinleyensis. A well
developed spots in males and, to some extent, in females is
another trait differing E. erynnin from E. ola.
Later Y. P. Korshunov (1996) rasied the status of the taxon
ola to a species and suggester the following improvement:
"HOLOTYPE: a male. F.w.l.: 25 mm. The wing upperside is evenly
soot-coloured. The underside is of the same colour but on the
fore wing, at the cell, there are groups of reddish scales
forming a hardly detectable sinking spot. On the hind wing
underside some scales are lighter than the neighbouring ones
where no bands are seen. In the genitalia the valva has a
rounded apex."
Erebia occulta sokhondinca Dubatolov et Zinchenko, 1995
Original description:
"Butterflies from the Chita Region, described below, are much
larger than E. o. occulta (f.w.l. being 19-21 mm) and
significantly larger than E. o. jakuta (f.w.l. 20-24 mm). They
well differ from both known subspecies by more developed fulvous
marking on the wings, often forming a contiguous band on the
fore wing. There is also a signofocant difference in the length
of the dentate part of the valva. Its ratio to the total length
of the valva in E. o. jacuta is 52-67% (with the mean 59,5%),
the samlpe size being 24, in new subspecies it is 56-67,4 (with
the mean 64%), the sample size being 20. The butterflies from
the East Sayan and Pribaikalye are close to the subspecies being
described.
Erebia occulta sokhondinca Dubatolov et Zinchenko, ssp.n.
Male: F.w.l. is 23 mm in the hlotype, 23-26 mm in paratypes.
The wing upperside is dark-brown, the fore wing bears 4 large
pale-fulvous spots along the outer margin, usually fused into a
band. They are centered with black dots elondate lengthwise with
respect to the wing. The hind wing has 3-4 round spots at the
outer margin, often centered with black dots. The pattern of the
fore wing underside is the same as on the upperside, but the fulvous
spots are always fused into a bright band. This band can extend
in direction of the wing base. The hind wing underside as in E.
o. jakuta.
Female: F.w.l. 23.5-26 mm. The pattern is as in males, but
the black dots are larger, up to 2 mm in diameter.
MATERIALS: The holotype - a male, 23.VI 1991, the Chita
Region, the Sokhondinskii nature reserve, the Verkhnii Bukukun
River (Dubatolov). Paratypes: 35 males 9 females, the same
locality and the mountain Tsagan-Ula, 19.VI - 6.VII 1991
(Dubatolov, Zinchenko and others)."
Erebia dabanensis olshvangi P. Gorbunov, 1995
Orininal description:
"..... The below described butterflies from the Polar Ural
differ from other subspecies by a darker brown-black ground
colour and the most substantial redction of brown postdiscal
spots....
Erebia dabanensis olshvangi P.Gorbunov, sbsp. n.
HOLOTYPE: a male. F.w.l.: 20 mm. Both sides of the fore wing
are dark-brown with four separate, diffuse on the underside,
fulvous-brown spots centered with small black dots; both sides
of the hind wing have three such spots; on the hind wing
underside the postdiscal area is to some extent lightened due to
greyish scales; the genitalia are analagous to those of other
subspecies.
ALLOTYPE: a female. F.w.l.: 20 mm. The wing upperside is
brown, the underside - greyish-brown; on the hind wing underside
a light-grey postdiscal band is clearly expressed; the fulvous-
brown pattern is similar with that of the holotype.
Generally the new subspecies is very variable. The length of
the fore wing ranges between 17 and 22 mm. In males the fulvous
spots on the upperside of both wings in 5-10 % of individuals
are absent. On the fore wing underside they, however, never
disappear entirely, on the contrary, in some specimens (15-20 %
of cases) they are fused into a common band, but its width, as
different from E. dabanensis troubridgei, as a rule does not
exceed 2 mm.
MATERIALS: the holotype: a male - 20.07.1992, Polar Ural, the
railway station Krasnyy Kamen', the mountain Slantsevaya, 300-
400 m above sea level, a montane lichen-fruticulose tundra
(P.Yu.Gorbunov leg.) The allotype - 10.07.1998, the same
locality (V.N.Olshvang leg.) Paratypes: 7 males 4 females - 10-
12.07.1998, the same locality; a male - 5.07.1990 - the same
locality; 4 males 3 females - 18-28.07.1992, the same locality.
The subspecies is named by the surname of Vladimir Nikolaevich
Olshvang, a researcher at IEPA UrD RAS [Institute of Ecology of
Plants and Animals of Ural Division of Russian Academy of
Sciences] , an entomologist who for a long time investigated
Polar Ural and collected a part of the type series."
Erebia theano tshugunovi Korshunov et Ivonin, 1995.
Original description:
"...Below we describe two subspecies from the Salairskii Kryazh
elevation and the Kuznetskii Alatau Mts. The former is
characterized by a brownish colour of spots on the wing
upperside. The butterflies from the Kuznetskii Alatau, West
Sayan and Tuva are characterized by a general reduction of
ochre-brown spots not forming contiguous bands.
Erebia theano tshugunovi Korshunov et Ivonin, sbsp. n.
HOLOTYPE: a male. F.w.l.: 18 mm. The wing upperside is dark-
brown; the postdiscal spots and the spot in the cell are brick-
red.The underside is the same as the upperside, but with a
greyish tint, which is more distinct along the wing margin; the
fulvous postdiscal spots on the hind wing consist of four hardly
noticeable dashes. The fringe is white with dark markings.
ALLOTYPE: a female. F.w.l.: 20 mm. The wing upperside as in the
male. The grey tint on the underside, especially on the hind
wing, is much more pronounced. The yellowish postdiscal spots on
the hind wing underside are smaller than on the fore wing (where
they are fulvous) and smaller than in the male; the basal spots
on the hind wing are small and hardly noticeable, as in the
male.
MATERIALS: the holotype: - 21.06.1994, the Novosibirsk Region,
the Kiternya river valley, a damp meadow (V.V. Ivonin); the
allotype: a female - 20.06.1994, the Novosibirsk region, the
Bol'shoy Elbash river valley, a damp meadow (V.V. Ivonin). The
paratypes: 11 males 8 females - 20.06.1994, the same locality; 5
males - 21.06.1994, the same locality, a female - 24.06.1994,
the same locality; 5 males - 21.06.1994, the Novosibirsk Region,
the village Novoselovo [this is a mistake - should be
Novososedovo, Yu.K.], a forest opening (V.V. Ionin); 1 male -
24.06.1949 - the Novokuznetsk city surroundings, the village
Torgay (A.E. Shtandel); 2 males - the Tomsk region, the village
Nizhnie Sokoly, a bogged meadow (G.S. Zolotarenko)
The subspecies is named in the honour of Sergey Mikhaylovich
Chugunov, a doctor of medicine at the Tomsk University, who in
1859-1920 comipled several annotated lists of Lepidoptera for
various regions of Siberia."
Erebia theano shoria Korshunov et Ivonin, 1995
Original description:
"Erebia theano shoria Korshunov et Ivonin, ssp. n.
HOLOTYPE: a male. F.w.l.: 19 mm. The wing upperside is brown
with the postdiscal bands composed of ochre-yellow spots and a
hardly noticeable spot in the cell. On the wing underside all
the spots are bright and large, including that in the cell of
the fore wing and the basal spots on the hind wing; the grey
suffusion is well expressed in the basal area. The fringe is
greyish, chequered.
ALLOTYPE: a female. F.w.l.: 19 mm. The wing upperside as in
the male, the spot in the cell is well expressed; on the wing
underside all the spots are bright and large; on the hind wing
the basal spots and grey suffusion are well expressed.
The specimens of the type series vary in the size of the lighter
spots, especially of the basal spots of the hid wing underside.
The fringe is brownish in some males.
MATERIALS: the holotype: a male - 5.07.1969, the Tuzukhsu river
valley in the Tom' river headwaters 1969 (Yu.P. Korshunov).
Allotype: a female - 13.07.1992, the Gornaya Shoria mountain
area, the Sheregesh settlement environs, the mountain Pustag,
1300 m [above sea level] (V.V. Ivonin). The paratypes: 21 males
- 5.07.1969, Gornaya Shoria, the Tuzukhsu valley; 13 males 9
females - 28.07.1969, the same locality; 86 males 28 females -
t24.06-27.07.1969, the Askiz River basin, Birikchul' (Yu.P.
Korshunov); 23 males - 17 and 24.07.1969, Khakassia, the Baza
river headwaters, (Yu.P. Korshunov); 4 males 2 females -
13.07.1992, the Sheregesh settlement, the mountain Pustag
(V.V.Ivonin)"
Erebia fletcheri chorymensis Korshunov, 1995.
Original description:
"...Below Erebia fletcheri chorymensis Korshunov, sbsp. n. is
described from the Kuznetskii Alatau Mts.
HOLOTYPE: a female. F.w.l.: 22 mm. Both wing sides are brown;
the fore wing has a wide fulvous-brown band, slightly tapering
to the anal angle, with four oval black spots about 1 mm long;
On the hind wing three [the words "of four" was erroneously
omited here - Yu.K.] fulvous spots contain black dots, as well
as an additional fifth small spot in front of the anal vein.
ALLOTYPE: a male. F.w.l.: 22 mm. The postdiscal band on
the fore wing upperside is darker and bears smaller black dots.
The paratypes have four of five fulvous spots on the hind
wing, of which one (in one specimen), three, four, or five being
centred with black dots. In the nominotypical subspecies (the
holotype being a female) the black spots on the fore wing are
round and almost twice as large, the second from the bottom being
almost thrice as large. Besides, in the butterflies from Altai
there is a tiny black spot at the apex which is not situated in
the same row with others, which is absent in all specimens from
the Kuznetskiy Alatau. The hind wing in the nominotypical
subspecies has four roundish reddish-fulvous spots, the two of
which at the anal angle being centered with black dots.
MATERIALS: the holotype: a female - 12.07.1975, the Kuznetskiy
Alatau, the environs of the mountain Bobrovaya (Yu.Korshunov);
the allotype: a male - the same locality and date; paratypes: 5
males 1 female - 9.07.1975, the same locality; 4 males 2 females
- 12.07.1975, the same locality; 4 males - 7.07.1975, the
Kuznetskii Alatau, the Sarala river valley; 4 males - 25-
26.06.1978, the same locality.
The name is derived from the Khakas word "khorym" - a
stone scree."
Satyrus ferula medvedevi Korshunov, 1996.
Original description:
"...The butterflies from Tuva, differing by bright and motled
pattern of the hind wing underside, with a contrasted band, are
dsacribed as Satyrus ferula medvedevi Korshunov, sbsp.n.
HOLOTYPE: a male. F.w.l. 27 mm. On the fore wing underside
the ocellus with a white pupil is almost four-fold larger than
the lower one (in altaica the ocelli are smaller and almost
of the same size). On the marbled hind wing underside the band
is rimmed with waving black lines, the line of the same
brightness and outline is conspicuous at the outer margin.
ALLOTYPE: A female. F.w.l. 26 mm. On the fore wing the large
black ocelli with white pupils are equal in size on both sides.
The hind wing underside ground colour is lighter than in males,
it is brownish-grey, with white scales along the outer margin of
the band and just at the wing margin. There are two black dots
at the anal angle, that about 2 mm in diameter and a much
smaller one.
MATERIALS: The holotype: a male - 19.07.1972, Tuva, Moren. The
allotype: a female - 23.07.1988, Tuva, the Kyzyl environs (V.
Zinchenko). Paratypes: 4 males - 18-30.07.1960, 40 km west of
[the village] Samagaltai, the surroundings of [the village]
Khol'-Ezhu, a grass/absinth and other type steppes; 2 males -
10.07.1961, Turan, Angol (A. Gukasyan); a male - 19.07.1972,
Moren; 2 males 6 females, 10-23.07.1990, 35 km NWW of
Samagaltai, the Shivilig-Khem basin (O. Kosterin).
The males are much more close to each other in appearance than
the females, among which there are individuals much darker above
but, on the contrary, lighter beneath than in the allotype.
The subspecies is dedicated to the memory of my teacher
Sergei Ivanovish Medvedev (1899-1979), who worked in Siberia
when was young.
...
Unfortunately, the comparative materials of a Chinese
subspecies liuplunschani O. Bang-Haas, 1993 is not available for
me."
Oeneis anna dzhulukuli Korshunov, 1998.
Original description:
"Elwes in the Altaian work of 1899 on Table 13 depicted three butterflies
as nanna. Of them two (fig. 4, a male, and fig. 8, a female) to our opinion,
just belong to the species anna. We have accumulated materials factually from
the same places where Elwes collected which allow to describe a new
subspecies dzhulukuli Korshunov, sbsp. n.
HOLOTYPE. A male. FWL 24 mm. Wing upperside brownish with an admixture
of ochre- and dark scales, that in general gives a pale-darkish background.
There are dark dots on outer area: on fore wing at apex a dot has a white
pupil, other dots below blind, in total four dots on fore wing, five blind
dots on hind wing. Androconial brands absent, no tufted scales found
on wing. Fringes chequered. Fore wing underside with a light-brown background
and four dots, as on upperside. On hind wing [upperside] there are many light
scales, veins white, among dots the upper one reduced so that only four
dots conspicuous. All the dots on both fore and hind wing [underside] with
white pupils. The genital apparatus is on the figure. In general it is of
the nanna type, differing by a somewhat different general configuration and
especially by valva outline.
ALLOTYPE. A female. FWL 24 mm. Wing upperside brownish, as in male, on hind
wing there is a darkening from base to outer field, black dots as in male.
And wing underside is similar
MATERIALS [in ISEA (Novosibirsk, Russia)]: Holotype - a male, 16.07.1982,
Taboshak Mt., Kuraiskii Mt. Range (Perunov, Y. E.). Allotype - a female,
31.07.1996, Dzhulukul’ Hollow, Bagayash, a damp floodland meadow,
2400 m (Bondarenko, A. V.). Paratypes - a female, 15.07.1978, Mukhor-Tarkhata
env., stow Argamzhi (A. Grazhdankin), a female, 16.07.1982, Taboshak Mt.
(Perunov, Y. E.) a female, 07.1982, Kuyakhtanar River - a Chuya River tributary
between Kurai and Kosh-Agach (exactly the place where Elwes collected!)
(Mertvetsv, D.), a female, 30.07.1985, v. Kalguty, a Poa/herbaceous
association (Malkov, Y. P.), a female and 2 males, 29-30.07.1996,
Dhulukul' Hollow, Bagayash, a lichen/dwarf-birch tundra (Bondarenko, A.V.)."
Oeneis jakutski Korshunov 1998.
Original description:
"HOPLOTYPE - a male. F.w.l. 22 mm. Wing upperside darkened from
bases to outer area, brownish, outer area lighter, ochre-brown, with three
small dots between veins on fore wing and four on hind wing. Androconial
scales noticeably grouped on fore wing at cell at spring of veins M2
and M3, while in nanna androconial scales spread along cell. Besides, vein
R5 in jakutski does not differ from others in thickness. The veins are
practically not swallen also in the paratypes. Fore wing underside greyish-
brown, cell with specks, its upper one third lighter, there is a brwn spot
near discal vein. Ocelli between veins (there are three of them) with white
pupils, slightly elongate, especially the upper one. Lighter scales form
somewhat like a halo around each ocellus. Apex marbled.
There are white streaks just between veins along wing margin (there are white
dots in nanna). On hind wing scales of medial band fuse with similar one at
wing base and on this continuous field only three small light spots are
noticeable just at fore margin. Band margin has no acute projections
(as different from nanna!), it is rimmed with whitish scales which alternate
with brown ones to form a marbled outer margin with four ocelli, from which
three are ocellate. Wing margin with small white streaks also but they are on
a background of other light scales. In the genitalia the differences from
nanna concern in the outline of the uncus, in longer subunci, in the outline
of the valca which has a peculiar projection on apex (see the figure).
ALLOTYPE - a female. F.w.l. 30 mm. Upperside of wings ochre-brown, their outers
areas lighter than other parts and bear an ocellus with pupil at apex and
two tiny dark spot below it at veins R5 and M3. A dark-brown spot on discal
vein is noticeable. Cell not darkened. Hind wing with four black spots on
outer margin bordered outside with a seen through band. Wing underside
marbled as specked with light and brownish scales, especially on hind
wing where very small ocelli with pupils are hardly seen among the scales.
On hind wing an ocellus with a pupil is conspicuous accompanied with a white
dot. Fringe chequered. The paratypes, although variable individualy, have
a smooth outline of the band margin. A presence of a triangular androconial
field on the male fore wing differ them from other species of the nanna-group.
MATERIAL - Holotype - a male. 9.06.1985, Yakutsk, botanical garden, a steppefied
bank slope of the Lena River, V. Dubatolov. ALlotype - a female, 16.06.1968,
Yakutsk, Chuchur-Muran. Paratypes: a male and female, 20.07.1962, Yakutsk, botanical
garden, Alexandrova, a male and female - 16.06.1968, Chuchur-Muran, a male -
19.06.1973, the Lena River valey, settlement Khaptagai, Yu. Ammosov, four
males - 10, 14, 15, 25.06.1974 - settlement Khaptagai, the Tamma River valley,
Yu. Ammosov, three males and a female - 1, 10, 17.06.1977 - 50 km S Yakutsk,
village Oktemtsy, Yu. Ammosov, a male - 9.06.1985, Yakutsk, botanical garden,
V. V. Dubatolov".
Oeneis sculda vadimi Korshunov, 1995
Original description:
"... Large and dark butterflies from N Pribaikalye are described
as Oeneis sculda vadimi Korshunov, sbsp.n, which, most
probably, ranges also in the northern E Siberia.
HOLOTYPE: a male. F.w.l.: 23 mm. The wing upperside is
bright pale-brown, the apex and the outer margin of the fore
wing are darkened, between the transversal vein and the apex
there is an ocellus with a pupil. The hind wing in the
submarginal area contains three black dots between the
veins. The wing underside is mottled due to a marble
pattern, the light veins are distinct, especially on the hind
wing. The postdiscal area of the fore wing contains
three, that of the hind wing - four ocelli with white pupils.
The discal band on the hind wing is dark-brown, with a
conspicuous rounded prominence behind the cell. the fringe
is chequered. In genitalia the shape of the uncus and the
valva processes and the structure of the aedeagus are somewhat
different from those in Oe. sculda sculda (see Appendix).
ALLOTYPE: a female. F.w.l.: 26 mm. The wing upperside is
brown, lighter than in the male; the fore wing has an ocellus
at the apex and another one at the anal angle; all the ocelli
are centered with white pupils; the wing underside resemble
that of the male as being equally bright; other characters are
also the same. In some paratypes the fore wings bear more
ocelli, there are females with the appearance indistinguishable
from the described male from above.
MATERIALS: the holtype: a male - 16.06.1988, the
Verkhneangarskiy mountain range, the surroundings of the town
Severobaikal'sk (V.V. Ivonin); the allotype: a female -
16.06.1988 - the same locality; paratypes - two females - 16
and 19.06.1988, the same locality.
The subspecies is named by the name of Vadim Vasilievich Ivonin
who participated on numerous expeditions in Siberia and the Far
East and collected the type series."
Oeneis elwesi ulugchemi Korshunov, 1995.
Original description:
"...According to observations in the Tuvinian Hollow (V.
Dubatolov, V. Zinchenko), the imagines keep to the stones,
hiding in their shade in a hot weather; they did not visit
flowers.
...The butterflies from the Tuvinian [intermontane]
Hollow and surrounding mountains differ by external characters
and also by habits. They are described as Oeneis elwesi
ulugchemi Korshunov, ssp. n.
HOLOTYPE: a male. F.w.l.: 23 mm. The wing upperside is brownish-
grey (darker than in ssp. tannuola), the outer margin of the
fore wing has light patches between the veins; the sex brand is
bright and occupies a part of the cell and some area ouside it,
there is a black oval spot at the fore wing apex. On the hind
wing a light postdiscal area, an oval black spot at the anal
angle, and a black dot above it are distinct. The fore wing
underside is ash-grey with a black oval spot at the apex; the
hind wing underside has a marble patern and a figured black
discal band (its shape differing from other subspecies) which is
fused with a dark scale suffusion at the wing base. The fringe
is white (in O. elwesi - chequered). The genital apparatus (see
Appendix) has a relatively long uncus and very uneven teeth on
the swallen end of the valva; in elwesi the valva apex has no
knobs and swellings, while the teeth are of relatively close
size. There is also a difference in the aedeagus structure. All
these details show that the rank of the taxon can be higher than
a subspecific.
ALLOTYPE: a female. F.w.l.: 24 mm. The internal half of the fore
wing upperside is brown, the external is sand-coloured with a
dark marginal rim; the postdiscal area contains three large oval
black spots; on the hind wing this area contains two black spots
(in other female paratypes they are reduced to dots or missing
at all). On the fore wing underside the black oval spots
present; the hind wing underside has a marble pattern, the area
between the band and margin being more heavily suffused with
dark scales than in the male.
The males other than holotype have 2-3 black spots on the fore
wing upperside (the upper is always larger) and always one
apical spot on the fore wing underside (sometmes black dots can
be seen beneath it). In some males the wings are rather dark,
but light areas are always retained, which sometimes contain
orange or even red-brown scales. The butterflies of the type
series are very variable in size (there are three among 29
females and ten among 110 males which are 1.5-2 times as small
as the holotype or allotype, respectively!).
MATERIALS: the holotype: a male - 10.05.1988, Tuva, the
surroundings of the Kyzyl city, the mountains at the Ulug-Khem
River right bank in the vicinity of the mouth of the Kaa-Khem
River (V. Zinchenko). The allotype: a female - 13.05.1989, the
same locality. Paratypes: 8 females - 13.05.1988, the same
locality; 1 male 1 female - 20.05.1989, the same locality; 94
males 26 females - 7-20.05.1990, the same locality (V.
Zinchenko, V. Dubatolov); a female - 10.05.1990, Tuva, the Ulug-
Khem district, 10 km east of the town Shagonar (V. Zinchenko)"
Oeneis magna pupavkini Korshunov, 1995
Original description:
"...In the Enisei River lower reaches and on the Polar Ural
butterflies occur with more clear-cut band on the wing
upperside, usually containing black spots, which are described
as Oeneis magna pupavkini Korshunov, sbsp.n.
HOLOTYPE: a male. F.w.l.: 26 mm. The fore wing upperside is
brown with a dark outer margin and a dark apex (that, as a rule,
is not met with in other subspecies), there are a black dot at
the apex and, below it, two smaller dots on fulvous spots; on
the hind wing [upperside] the fulvous band occupies its outer
part almost entirely, it contains two black dots at the anal
angle. The fringe is chequered, black-and-white. There are light
scales on the fore wing underside which hide the dark marginal
rim, yellow spots and black dots; the hind wing underside has a
marble pattern, the light scales concentrating at the outer
margin of the discal band and at the wing base above the cell.
Other males have the same black dots or lack them, while one
of them has large black spots, as in a female, which it was
considered to be at the first glance.
ALLOTYPE: a female. F.w.l.: 27 mm. The wing upperside is
brighter than in the male, the fulvous spots, containing black
dots, on the fore wing are fused into a band. The black spots
are large and centered with white pupils on both wing sides; the
fore wing apex has a marble pattern underside; the hind wing
underside is brighter than in the male.
MATERIALS: the holotype: a male - 19.07.1978, the Putorana
Plateau, the middle reaches of the Rybnaya river (D.I.Pupavkin).
The allotype: a female - 12.07.1978, the same locality, a larch
taiga. Paratypes: 1 male 1 female - 7-12.07.1978, the same
locality, forest-tundra; 12 males 2 females - 15-27.07.1978,
the same locality; a female - 1.08.1973, the Polar Ural, [the
railway station] Kharp (V.N. Olshvang); 1 male 4 females - 4-
10.07.1994, the same locality (P. Gorbunov).
The subspecies is named and dedicated to the
memory of Dmitriy Maksomovich Pupavkin, a forest entomologist
and an explorer of SIberia in 70s-80s years. "
Oeneis aktashi ona Korshunov, 1996.
Original description:
"...The butterflies from the W Sayan are different by a
greater developmemnt of pale-ochre coloured spots in the wing
submarginal area, an often presence of dot ocelli, by brighter
coloured females. Below we give a description of Oeneis aktashi
ona Korshunov, sbsp.n.
HOLOTYPE: a male. F.w.l. 24 mm. The wing upperside is
brownish with pale-ochre brands along the outer margin,
especially on the hind wing. There is a dark dot at the fore
wing apex. The fore wing underside is evenly brown being lighter
than the hind wing, which has a marble pattern and is darker in
the area from the base to the cell apex, a band is noticeable
with more or less rounded margin (which is, by the way, angular
in O. sarala).
ALLOTYPE: a female. F.w.l. 23 mm. The upperside of wings is
brown, their outer parts are ochre-brown owing to ochre-coloured
scales forming a distinct field on the fore wing with two dark
dots on it. The hind wing upperside is mottled due to alteration
of dark, brown and ochre scales. On the wing underside the same
alteration makes a marble pattern, especially on the hind wing.
Only one dark dot is seen on the fore wing apex.
MATERIALS: The holotype: a male - 28.07.1975. The Ona River
headwaters, the Sayan Pass, the Saylyg-Khem-Taiga mountain
range, a detritous ridge with Dryas, Betula nana rotundifolia,
snow patches (Yu. P. Korshunov). The allotype: a female -
25.07.1975, the same locality. Paratypes: 15 males 9 females,
25-28.07.1975, the same locality; 1 male - 23.07.1993, the
mountain Mungun-Taiga, barren summits (D.V. Logunov).
In the paratypes the dark dot on the fore wing is seen in 4
specimens of 15. In almost a half of the paratypes the ochre
spots are so much developed that the brown ground colour retains
in the wing outer part as a narrow marginal band. In some
females the mottleness is weakened, especially above, in some of
them only one or no dark dot at the apex is seen. In some
females the marble pattern beneath is paler. The specimen of
1993 is peculiar - it is almost twice as small and the outer
fielw on the hind wing underside is much lightened."
Oeneis chione tshukota Korshunov 1998.
Original description:
"HOLOTYPE: A male - F.w.l. 23 mm. Upperside light-brown, on fore wing below
cell a black androconial stroke is well seen. On wing outer area there is
a black spot at apex and between veins Cu and M. On hind wing there is a
black spot between cubital veins. Dots have white pupils both above and beneath.
Fore wing underside light-brown with a marbled apex, hind wing with a band
somewhat resembling that in dzhugdzhurim surrounded with light scales which
cover the most part of cell. Male genital apparatus see on Fig. It is of a
polyxenes type, differs by a peculiar structure of the uncus and valva.
ALLOTYPE. Female - F.w.l. 26 mm. Wing underside light-brown, sandy, on fore
wing there are four black dots, one of which being tiny and the upper one with
a white pupil; on hind wing above two large black dots in anal angle and
a small one above. On hind wing underside dots as if darkened with brown scales,
band wider than in male, cell bears more brown scales than light ones.
Fringe chequered in males and females.
MATERIAL. Holotype - a male, 24.06.1967, 20 km of Markovo upstream the
Anadyr' River (A. Mirzaeva). Allotype - a female, 27.06.1967, 5 km W of
Markovo a meadow at a birch grove. Paratypes - two males - 26.06 and
10.07.1967, the same locality as the holotype, Poaceae/Cyperaceae community.
Seven males and seven females, 24, 26, 27.06.1967, the same locality as
the allotype (V. Skutov).
COMMENT. The butterflies to some extent resemble the western norna, differing
in the pattern, fringe, detail of the genital apparatus. As a matter of fact,
there is no full similarity between the specimens of tshukota. Among the paratypes
the male has only a dot on either wing and a band intermediate between the
holotype and allotype; in females there is a difference in black dots: from
one to two on fore wing to four on the left wing and three on the right wing.
The bands are more or less similar but the suffusion is of different proportion]
of light and brown scales."
Lycaena phlaeas ganalica P. Gorbunov, 1995.
Original description:
" ...Peculiar butterflies from Kamchatka are described as
Lycaena phlaeas ganalica P.Gorbunov, sbsp. n. MALE: F.w.l. is
13.8 mm in the holotype, 14.1 in the paratype. The fore wing
upperside in the basal, discal and postdiscal areas is copper-
coloured with suffusion of dark scales and eight black spots, in
general it is darker than in other Siberian subspecies. The
brown-grey margin is about 2 mm wide. The fore wing apex seems
to be more acute and the angle of the outer and anal margins
more blunt than in the subspecies phlaeas and polaris. The hind
wing upperside is ash-grey with, an orange streak 1.3-1.5 mm
wide, with waving outer margin, along the wing outer margin. The
underside is light-grey, in the central area of the fore wing it
is yellowish-orange, generally noticeablylighter than in other
Siberian subspecies. The pattern of black spot repeats that of
phlaeas and polaris. The orange submarginal spots are almost
invisible in the holotype and are represented by a fragmented
streak, containing of five strokes, indented to 1.5-2 mm of the
wing outer margin.
MATERIALS: the holotype: a male - 14.07.83, Kamchatka,
Ganal'skaya Tundra, (V.N.Olshvang). A paratype: a male -
14.07.83 the same locality."
Note: To my mind, this is not a subspecies but just one
more variety of this greatly variable species. There are
specimens from Kamchatka in the ISEA collection quite different
from the described one.
Heodes alciphron rubidus Korshunov, 1995
The name was proposed in Korshunov & Gorbunov (1995) as
follows: "...A male from Mongolia described by Yu.P. Kosrhunov
(1977) is designated here as Heodes alciphron rubidus Korsunov,
sbsp. n. Additional material: a female - 16.07.1939, the Chita
River headwaters (E.I. Pavlova)". Translation of the cited
description, which should be considered as the original
description of the subspecies, is follwoing:
"Central [Aimak]: the stow Sudzukte, 15-16 VII 1925, 1 male
[(P.K. Kozlov, E.V. Kozlova and colleagues)].
MALE: The fore wing length is 16 mm. The violet flash on the
wings is more weak expressed than in the typical form (the
butterfly looks reddish), while the dark spots on the fore wing
are more contrasted, especially in the cell. Almost double spots
at the hind margin of the fore wing are well seen. On the hind
wing underside the dark spots of the second row internally of
the orange streak are not round, as in the typical form, but
somewhat elongate, they are not ringed with light rims. The
genitalia scarcely differ from the type."
Plebejus argus caerulea P. Gorbunov, 1995.
Original description:
"... Blue upperside ground colour is found also in the males of
P. argus from Central Yakutia. They approach by a number of
external features to Plebejus tancrei verknojanicus, known from
Yakutia, but well differ from them by the genitalia (see
Appendix). We describe them here as Plebejus argus caerulea P.
Gorbunov, sbsp. n.
HOLOTYPE: - a male. F.w.l.: 12.5 mm. The wing upperside is
glittering-blue, whereas violet in other Siberian subspeceis.
The dark border is about 1 mm wide, on the hind wing it is
contacted by blask spots situated between the veins. The wing
underside ground colour is pale-grey, a bit darker than in the
South Yakutian butterflies of ssp. clarasiatica. On the fore
wing upperside the marginal spots are small, clear-cut, the row
of the postdiscal spots is less curved than in ssp.
clarasiatica.
MATERIALS: the holotype: a male - 28.06.1992, Central Yakutia,
the Yakutsk city, the Botany Garden, a meadow on a larch wood
edge (P. Gorbunov). Paratypes: a male - 28.06.1992, the same
locality; 2 males - 2.07.1968, Yakutia (Zakharov)"
Note: in fact this taxon resulted from confsion of the genitalia
preparations, the characters described belonged to a specimen of
Plebejus tancrei (P. Gorbunov, personal communication).
Aricia eumedon fylgides P. Gorbunov, 1995.
Original description:
"...All the specimens from Kamchatka, known to us, have no
white beam on the hind wing underside and are characterized by
the black dots and orange spots (in males) of the hind wing
underside smaller than in typical butterflies. We describe them
as a subspecies Aricia eumedon fylgides P. Gorbunov, sbsp.n.
MALE. F.w.l. 11.2-13.0 mm. The wing upperside is evenly dark-
brown. The wing underside ground colour is ash-grey. Five
postdiscal spots on the fore wing and six on the hind wing are
significantly smaller than in the nominotypical subspecies. The
black discal spot on the fore wing is as well smaller. The
discal spot on the hind wing is white, triangular in shape, in
two paratypes with black scales in the centre. The white beam is
absent. Dark spots at the fore wing outer margin are reduced to
traces, the orange spots (peculiar to the subspecies eumedon)
are not seen here. On the fore wing underside the orange spots
are smallet than in the subspecies eumedon, their size reducing
from the anal angle to the apex, the dark lulunes accompanying
they internaly are weakly expressed. The hind wing base has a
suffusion of glittering blueish scales, the black basal spots
are absent or only one of them, closest to the fore margin, is
present. The fringe is evenly white.
FEMALE. F.w.l. 13.4-14.2 mm. The wing upperside is brown with
three orange lunules at the anal angle of the hind wing. The
wing underside ground colour is brown-grey. The basal. discal
and the postdiscal black spots are as in males. The orange
submarginal spots are much larger, 3-4 of such pale-orange spots
are present also on the fore wing. The black lunules
(internally) and black dots (externally) accompanying the orange
spots are less expressed than in the nominotypical subspecies.
MATERIALS: The holotype: a male - 22.07.1983, Kamchatka,
Paratunka (V.N. Olshvang). Paratypes: 2 males - 22.07.1983, the
same locality; 3 males 1 female - 14.07.1983, Kamchatka,
ganal'skaya Tundra (V.N. Olshvang); 3 males 1 female -
6.07.1968, Kamchatka, the Mil'kovo District, the middle flow of
the Kavycha River; 4 males - 18.06.1984, Kamchatka, the environs
of the city of Petropavlovsk."
Polyommatus icarus korshunovi P. Gorbunov, 1995
Original description:
"In the populations of S Siberia the appearance of
butterflies is very diverse. However, in arid regions
in average smaller individuals, with somewhat narrower wings, a
whitish wing underside ground colour in males and greyish in
females, evidently predominate. The orange submarginal spots in
them are smaller than in the butterflies from more northern
regions and are isolated from each other. The females have or
have not red small submarginal lunules on the wing upperside,
which is often substantially suffused with blue scales. Based on
these characters we state a subspecies Polyommatus icarus
korshunovi P. Gorbunov, sbsp.n.
MATERIALS: The holotype: a male - 26.05.1990, Tuva, the Erzin
River valley (V.V. Dubatolov). Paratypes: a male - the same
locality; 2 females - 27.07.1972, Tuva, Toora-Khem (Yu.P.
Korshunov); a female - 19.08.1962, Tuva, the Tes[-Khem] River
(L. Violovich); a fenale and male - 17.06.1963, the same
locality; 2 males - 18.06, 1987, Tuva, the Toszha District, Lake
Azas, (V. Zinchenko); a male - 1-5.05.1993, Tuva, the Kyzyl city
environs (D. Logunov); 2 males - Tuva, the Shivilig-Khem River,
a bushy steppe (O. Kosterin); 3 males - 21.06.1907, SE Altai,
the Chuiskaya Steppe, Kosh-Agach; a female - 13.07.1097, the
Chuiskii Tract road, the Kuraiskaya Steppe [a collector is
dropped - it was E.G. Rodd] ; a female - 19.07.1966, SE ALtai,
between the mountain Supor and the Chuiskaya Steppe, 2000-2400 m
[above the sea level].
Polyommatus icarus omelkoi Dubatolov et Korshunov, 1995
Original description:
"Butterflies from the southern Far East differ from the
Siberian ones by in average larger size (f.w.l. being 17-19 mm
in males, 16.5-18 mm), reduction of the basal suffusion with
glittering scales on the hind wing underside, which usually does
not extend out of the row of basal spots, enlarged black
dots on the wing underside, and a lighter (whitish) wing
underside ground colour in males. By these characters we state a
subspecies Polyommatus icarus omelkoi Dubatolov et Korshunov,
sbsp.n.
MATERIALS: The holotype: a male - 17.07.1993, Primorie, the
Anuchinskii District, 14 km north of Chernyshevka (V.V.
Dubatolov, V.K. Zinchenko). Paratypes: 2 males 2 females -
15.07.1993, the same locality; 1 male - 18.07.1993, 20 km NNW of
Chernyshevka, Kamenistyi Klyuch, a road in a montane broad-
leaved forest; 4 males - 11 and 13.07.1981, 21.07.1982,
11.07.1985, N Korea (Im Khon An).
The subspeceis is named after the surname of Omel'ko, Michail
Michailovich, a lepidopterologist at the Ussuriiskii
Reservation, an explorer of the life history of a number of
butterfly species of S Primorie."
Names suggested to replace junior homonyms.
The name kurentzovi Korshunov, 1995 has been suggested for
the subspecies Pyrgus centaureae sibirica Kurenzov, 1970
(ranging in the Magadan Region and the Koryak Upland and
characterized by a darker ground colour of the wing upperside
and a reduced white spots on the hind wing upperside), since
sibirica Kurenzov is a secondary homonym of Pyrgus sibirica
(Reverdin, 1911).
The name Colias nastes dezhnevi Korshunov, 1995 has been
suggested to replace Colias nastes sibirica Kurenzov, 1970,
since the latter was preoccupied by Colias aurora sibirica
Lederer, 1852. This subspecies, reported for Chukotka and the
Kolyma River basin, has a pale greenish-yellow wing upperside
ground colour, the light spots on the hind wing margin are of
the ground colour.
The name sachelininsis in Clossiana oscarus sachalinensis
Matsumura, 1936 was replaced, as an homonym, by the name
insularia Korshunov, 1996.
The trinomen Erebia euryale iremelica Korshunov, 1995 was
proposed to replace Erebia euryale uralensis Goltz, 1930 (=
euryaloides auct., nec Tngstrom, 1869) which is a junior homonym
for Erebia aethiops uralensis. This subspecies differ from the
typical butterflies of Central Europe by small blind ocelli,
which reside ona wide brownish-red band on the fore wing and
often absent on the hind wing.
The subspecies Erebia pandrose orientalis Goltz, 1930, was
described for the mountains of S. Siberia, but this name
was preoccupated by Erebia epiphron orientalis Elwes, 1900. For
this reason I have replaced this name by yernikensis Korshunov, 1994,
nom. nov. It differs from the butterflies of N Europe by in
general smaller size and more faded colour of the postdiscal
field on the fore wing upperside.
Pseudochazara tshujaca Korshunov, 1998 (=pallida auct.)
Translation of systematic notes:
"The form pallida Staudingerm 1901 was stated from specimens from
Spain (Granada, Andalusia) and by Elwes' Altaian collections.
As still N. J. Kusnezov (1909) has noted, the name pallida is
useful for designation of either small and pale butterflies. In
this situation it is correct for us to introduce the enw name in
a hope that someone who would designate a lectotype of pallida
would choose a Spanish specimen and would not attempt to close
this our initiative, as it recently took place with lederi and
close taxa.
The species tshujaca appeared an endemic of Altai, in this respect
it resembles a Crimean euxina. While in Crimea an island isolation
explains much, in Altai the reasons were perhaps more complicated,
perhaps gene drift or a special reaction to extremum highland
condition. The male genital apparatus is slightly affected, there is
no specific differences from hippolyte, as well as in euxina. V. Ivonin
has noticed that in tshujaka and hippolyte, except for the difference
in the visible androconial spots, there is a difference in the androconial
scales and as well on disposition and colour of scales on the antennal
club: in hyppolyte the entire club is scale-clad while in tshujaca
only in part."
A name otton Korshunov, 1996 was proposed for a subspecies
of Plebejus lucifer instead of biton Bremer, 1861, which is a
primary homonym to biton Sulz, 1776 - a subspecies of
Polyommatus damon (D. et Sch., 1775).
A name Aricia nicias kolosovi Korshunov, 1995 was proposed
pro Lycaena nicias septentrionalis Krulikowsky in Seitz, 1909,
nec. Lycaena icarus septentrionalis Fusch, 1900, differing from
the Central European butterflies by a paler wing underside and
in average narrower marginal band in males.
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Korshunov, Yu. P. 1977. [Butterflies (Lepidoptera,
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Korshunov, Yu. P. 1985. [Notes on the systematics of the
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